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Stock-based Compensation (Tables)

9 Months Ended

Sep. 30, 2022

Stock-based Compensation

Schedule of stock-based compensation expense

Total compensation cost recognized for all stock-based compensation awards in the condensed consolidated statements of operations was as follows (in thousands):

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Three Months Ended

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Nine Months Ended

​

​

September 30,

​

September 30,

​

​

2022

​

2021

​

2022

​

2021

Research and development

​

$

3,045

​

$

4,552

​

$

9,803

​

$

12,689

General and administrative

​

2,836

​

5,460

​

14,127

​

23,053

Total stock-based compensation expense

​

$

5,881

​

$

10,012

​

$

23,930

​

$

35,742

Schedule of changes in unvested restricted stock

The following is a summary of restricted stock and restricted stock unit awards activity for the nine months ended September 30, 2022:

​	​	​	​	​	​
​		​		Weighted
​	​	​	​	Average
​	​	​	​	Grant Date
​	​	​	​	Fair Value
​	​	Shares	​	Per Share
Unvested restricted stock and restricted stock unit awards as of December 31, 2021		628,732	​	$	41.28
Issued		1,334,826	​	$	15.57
Vested		(271,037)	​	$	44.69
Forfeited	​	(188,839)	​	$	31.88
Unvested restricted stock and restricted stock unit awards as of September 30, 2022		1,503,682	​	$	19.02

Schedule of stock option activity

The following is a summary of stock option activity for the nine months ended September 30, 2022:

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​	​	​	​	​	​	​	​	​	​	​
​		​		Weighted Average			Remaining		Aggregate Intrinsic
​	​	Shares	​	Exercise Price		​	Contractual Life (years)	​	Value (in thousands)
Outstanding at December 31, 2021		3,016,085	​	$	34.24	​	7.5	​	$	5,052,469
Granted	​	2,749,534	​	$	14.64	​	​	​	​	​
Exercised	​	(17,569)	​	$	16.97	​	​	​	​	​
Cancelled	​	(531,926)	​	$	30.78	​	​	​	​	​
Outstanding at September 30, 2022		5,216,124	​	$	24.32	​	8.2	​	$	1,534,373
Exercisable at September 30, 2022		1,906,975	​	$	30.80	​	6.8	​	$	807,066

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Tabular disclosure of the changes in outstanding nonvested restricted stock shares.

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