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Condensed Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Other Comprehensive (Loss) Gain | Other Accumulated Deficit
--- | --- | --- | --- | --- | ---
Balance, beginning of period (in shares) at Dec. 31, 2021 |  | 68,435,257 |  |  |
Balance, beginning of period at Dec. 31, 2021 | $ 553,642 | $ 7 | $ 1,411,827 | $ (493) | $ (857,699)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Exercise of stock options (in shares) |  | 12,573 |  |  |
Exercise of stock options | 218 |  | 218 |  |
Stock-based compensation expense | 11,431 |  | 11,431 |  |
Vesting of restricted common stock awards (in shares) |  | 154,834 |  |  |
Vesting of restricted common stock awards | 0 |  |  |  |
Unrealized gain (loss) on marketable debt securities | (2,016) |  |  | (2,016) |
Net loss | (50,515) |  |  |  | (50,515)
Balance, end of period (in shares) at Mar. 31, 2022 |  | 68,602,664 |  |  |
Balance, end of period at Mar. 31, 2022 | 512,760 | $ 7 | 1,423,476 | (2,509) | (908,214)
Balance, beginning of period (in shares) at Dec. 31, 2021 |  | 68,435,257 |  |  |
Balance, beginning of period at Dec. 31, 2021 | 553,642 | $ 7 | 1,411,827 | (493) | (857,699)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Unrealized gain (loss) on marketable debt securities | (3,798) |  |  |  |
Net loss | (159,694) |  |  |  |
Balance, end of period (in shares) at Sep. 30, 2022 |  | 68,761,729 |  |  |
Balance, end of period at Sep. 30, 2022 | 414,745 | $ 7 | 1,436,422 | (4,291) | (1,017,393)
Balance, beginning of period (in shares) at Mar. 31, 2022 |  | 68,602,664 |  |  |
Balance, beginning of period at Mar. 31, 2022 | 512,760 | $ 7 | 1,423,476 | (2,509) | (908,214)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Exercise of stock options (in shares) |  | 20 |  |  |
Exercise of stock options | 0 |  |  |  |
Stock-based compensation expense | 6,618 |  | 6,618 |  |
Vesting of restricted common stock awards (in shares) |  | 77,884 |  |  |
Vesting of restricted common stock awards | 0 |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  | 37,866 |  |  |
Issuance of common stock under employee stock purchase plan | 367 |  | 367 |  |
Unrealized gain (loss) on marketable debt securities | (878) |  |  | (878) |
Net loss | (53,453) |  |  |  | (53,453)
Balance, end of period (in shares) at Jun. 30, 2022 |  | 68,718,434 |  |  |
Balance, end of period at Jun. 30, 2022 | 465,414 | $ 7 | 1,430,461 | (3,387) | (961,667)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Exercise of stock options (in shares) |  | 4,976 |  |  |
Exercise of stock options | 80 |  | 80 |  |
Stock-based compensation expense | 5,881 |  | 5,881 |  |
Vesting of restricted common stock awards (in shares) |  | 38,319 |  |  |
Vesting of restricted common stock awards | 0 |  |  |  |
Unrealized gain (loss) on marketable debt securities | (904) |  |  | (904) |
Net loss | (55,726) |  |  |  | (55,726)
Balance, end of period (in shares) at Sep. 30, 2022 |  | 68,761,729 |  |  |
Balance, end of period at Sep. 30, 2022 | 414,745 | $ 7 | 1,436,422 | (4,291) | (1,017,393)
Balance, beginning of period (in shares) at Dec. 31, 2022 |  | 68,847,382 |  |  |
Balance, beginning of period at Dec. 31, 2022 | 360,680 | $ 7 | 1,442,405 | (3,601) | (1,078,131)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Stock-based compensation expense | 4,507 |  | 4,507 |  |
Vesting of restricted common stock awards (in shares) |  | 146,209 |  |  |
Vesting of restricted common stock awards | 0 |  |  |  |
Unrealized gain (loss) on marketable debt securities | 1,322 |  |  | 1,322 |
Net loss | (49,036) |  |  |  | (49,036)
Balance, end of period (in shares) at Mar. 31, 2023 |  | 68,993,591 |  |  |
Balance, end of period at Mar. 31, 2023 | 317,473 | $ 7 | 1,446,912 | (2,279) | (1,127,167)
Balance, beginning of period (in shares) at Dec. 31, 2022 |  | 68,847,382 |  |  |
Balance, beginning of period at Dec. 31, 2022 | 360,680 | $ 7 | 1,442,405 | (3,601) | (1,078,131)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Unrealized gain (loss) on marketable debt securities | 2,149 |  |  |  |
Net loss | (134,345) |  |  |  |
Balance, end of period (in shares) at Sep. 30, 2023 |  | 81,668,796 |  |  |
Balance, end of period at Sep. 30, 2023 | 360,462 | $ 8 | 1,574,382 | (1,452) | (1,212,476)
Balance, beginning of period (in shares) at Mar. 31, 2023 |  | 68,993,591 |  |  |
Balance, beginning of period at Mar. 31, 2023 | 317,473 | $ 7 | 1,446,912 | (2,279) | (1,127,167)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Issuance of common stock from public offering, net commissions, underwriting discounts and offering costs (in shares) |  | 12,500,000 |  |  |
Issuance of common stock from public offering, net commissions, underwriting discounts and offering costs | 117,079 | $ 1 | 117,078 |  |
Exercise of stock options (in shares) |  | 3,122 |  |  |
Exercise of stock options | 11 |  | 11 |  |
Stock-based compensation expense | 5,215 |  | 5,215 |  |
Vesting of restricted common stock awards (in shares) |  | 64,492 |  |  |
Vesting of restricted common stock awards | 0 |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  | 55,704 |  |  |
Issuance of common stock under employee stock purchase plan | 435 |  | 435 |  |
Unrealized gain (loss) on marketable debt securities | (6) |  |  | (6) |
Net loss | (40,290) |  |  |  | (40,290)
Balance, end of period (in shares) at Jun. 30, 2023 |  | 81,616,909 |  |  |
Balance, end of period at Jun. 30, 2023 | 399,917 | $ 8 | 1,569,651 | (2,285) | (1,167,457)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Stock-based compensation expense | 4,731 |  | 4,731 |  |
Vesting of restricted common stock awards (in shares) |  | 51,887 |  |  |
Vesting of restricted common stock awards | 0 |  |  |  |
Unrealized gain (loss) on marketable debt securities | 833 |  |  | 833 |
Net loss | (45,019) |  |  |  | (45,019)
Balance, end of period (in shares) at Sep. 30, 2023 |  | 81,668,796 |  |  |
Balance, end of period at Sep. 30, 2023 | $ 360,462 | $ 8 | $ 1,574,382 | $ (1,452) | $ (1,212,476)

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