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Condensed Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Other Comprehensive Income (Loss) | Accumulated Deficit
--- | --- | --- | --- | --- | ---
Balance, beginning of period (in shares) at Dec. 31, 2023 |  | 81,767,263 |  |  |
Balance, beginning of period at Dec. 31, 2023 | $ 349,097 | $ 8 | $ 1,580,241 | $ 198 | $ (1,231,350)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Exercise of stock options (in shares) |  | 21,975 |  |  |
Exercise of stock options | 192 |  | 192 |  |
Vesting of restricted common stock awards (in shares) |  | 445,713 |  |  |
Stock-based compensation expense | 7,585 |  | 7,585 |  |
Unrealized (loss) gain on marketable debt securities | (524) |  |  | (524) |
Net loss | (61,950) |  |  |  | (61,950)
Balance, end of period (in shares) at Mar. 31, 2024 |  | 82,234,951 |  |  |
Balance, end of period at Mar. 31, 2024 | 294,400 | $ 8 | 1,588,018 | (326) | (1,293,300)
Balance, beginning of period (in shares) at Dec. 31, 2023 |  | 81,767,263 |  |  |
Balance, beginning of period at Dec. 31, 2023 | 349,097 | $ 8 | 1,580,241 | 198 | (1,231,350)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Unrealized (loss) gain on marketable debt securities | 324 |  |  |  |
Net loss | (191,698) |  |  |  |
Balance, end of period (in shares) at Sep. 30, 2024 |  | 82,538,624 |  |  |
Balance, end of period at Sep. 30, 2024 | 175,634 | $ 8 | 1,598,152 | 522 | (1,423,048)
Balance, beginning of period (in shares) at Mar. 31, 2024 |  | 82,234,951 |  |  |
Balance, beginning of period at Mar. 31, 2024 | 294,400 | $ 8 | 1,588,018 | (326) | (1,293,300)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Vesting of restricted common stock awards (in shares) |  | 121,643 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  | 72,920 |  |  |
Issuance of common stock under employee stock purchase plan | 322 |  | 322 |  |
Stock-based compensation expense | 5,010 |  | 5,010 |  |
Unrealized (loss) gain on marketable debt securities | (116) |  |  | (116) |
Net loss | (67,607) |  |  |  | (67,607)
Balance, end of period (in shares) at Jun. 30, 2024 |  | 82,429,514 |  |  |
Balance, end of period at Jun. 30, 2024 | 232,009 | $ 8 | 1,593,350 | (442) | (1,360,907)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Vesting of restricted common stock awards (in shares) |  | 109,110 |  |  |
Stock-based compensation expense | 4,802 |  | 4,802 |  |
Unrealized (loss) gain on marketable debt securities | 964 |  |  | 964 |
Net loss | (62,141) |  |  |  | (62,141)
Balance, end of period (in shares) at Sep. 30, 2024 |  | 82,538,624 |  |  |
Balance, end of period at Sep. 30, 2024 | 175,634 | $ 8 | 1,598,152 | 522 | (1,423,048)
Balance, beginning of period (in shares) at Dec. 31, 2024 |  | 82,734,696 |  |  |
Balance, beginning of period at Dec. 31, 2024 | 134,274 | $ 8 | 1,602,441 | 268 | (1,468,443)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Issuance of common stock from at the market equity offering, net (in shares) |  | 706,236 |  |  |
Issuance of common stock from at-the-market equity offering, net | 1,435 |  | 1,435 |  |
Vesting of restricted common stock awards (in shares) |  | 268,604 |  |  |
Stock-based compensation expense | 2,979 |  | 2,979 |  |
Unrealized (loss) gain on marketable debt securities | (180) |  |  | (180) |
Net loss | (76,088) |  |  |  | (76,088)
Balance, end of period (in shares) at Mar. 31, 2025 |  | 83,709,536 |  |  |
Balance, end of period at Mar. 31, 2025 | 62,420 | $ 8 | 1,606,855 | 88 | (1,544,531)
Balance, beginning of period (in shares) at Dec. 31, 2024 |  | 82,734,696 |  |  |
Balance, beginning of period at Dec. 31, 2024 | 134,274 | $ 8 | 1,602,441 | 268 | (1,468,443)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Unrealized (loss) gain on marketable debt securities | (268) |  |  |  |
Net loss | (154,440) |  |  |  |
Balance, end of period (in shares) at Sep. 30, 2025 |  | 93,264,421 |  |  |
Balance, end of period at Sep. 30, 2025 | 13,453 | $ 9 | 1,636,327 | 0 | (1,622,883)
Balance, beginning of period (in shares) at Mar. 31, 2025 |  | 83,709,536 |  |  |
Balance, beginning of period at Mar. 31, 2025 | 62,420 | $ 8 | 1,606,855 | 88 | (1,544,531)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Issuance of common stock from at the market equity offering, net (in shares) |  | 3,419,836 |  |  |
Issuance of common stock from at-the-market equity offering, net | 7,185 | $ 1 | 7,184 |  |
Vesting of restricted common stock awards (in shares) |  | 102,894 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  | 156,159 |  |  |
Issuance of common stock under employee stock purchase plan | 228 |  | 228 |  |
Stock-based compensation expense | 2,664 |  | 2,664 |  |
Unrealized (loss) gain on marketable debt securities | (73) |  |  | (73) |
Net loss | (53,235) |  |  |  | (53,235)
Balance, end of period (in shares) at Jun. 30, 2025 |  | 87,388,425 |  |  |
Balance, end of period at Jun. 30, 2025 | 19,189 | $ 9 | 1,616,931 | 15 | (1,597,766)
Statement of Stockholders' Equity (Deficit) |  |  |  |  |
Issuance of common stock from at the market equity offering, net (in shares) |  | 5,799,178 |  |  |
Issuance of common stock from at-the-market equity offering, net | 17,147 |  | 17,147 |  |
Exercise of stock options (in shares) |  | 12,130 |  |  |
Exercise of stock options | 21 |  | 21 |  |
Vesting of restricted common stock awards (in shares) |  | 64,688 |  |  |
Stock-based compensation expense | 2,228 |  | 2,228 |  |
Unrealized (loss) gain on marketable debt securities | (15) |  |  | (15) |
Net loss | (25,117) |  |  |  | (25,117)
Balance, end of period (in shares) at Sep. 30, 2025 |  | 93,264,421 |  |  |
Balance, end of period at Sep. 30, 2025 | $ 13,453 | $ 9 | $ 1,636,327 | $ 0 | $ (1,622,883)

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