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CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
shares in Thousands, $ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Other Comprehensive Income (Loss)
Accumulated Deficit
Balance at Dec. 31, 2015 $ 94,765 $ 99 $ 685,189 $ 7,289 $ (597,812)
Balance (in shares) at Dec. 31, 2015   99,095      
Net loss (62,906)       (62,906)
Other comprehensive loss (7,186)     (7,186)  
Issuance of common stock from public offering, net of offering costs 21,981 $ 3 21,978    
Issuance of common stock from public offering, net of offering costs (in shares)   3,526      
Issuance of common stock from exercise of stock options and purchases from ESPP 3,068 $ 1 3,067    
Issuance of common stock from exercise of stock options and purchases from ESPP (In Shares)   854      
Stock-based compensation 8,065   8,065    
Balance at Dec. 31, 2016 57,787 $ 103 718,299 103 (660,718)
Balance (in shares) at Dec. 31, 2016   103,475      
Net loss (60,585)       (60,585)
Other comprehensive loss (200)     (200)  
Issuance of common stock from public offering, net of offering costs 30,156 $ 11 30,145    
Issuance of common stock from public offering, net of offering costs (in shares)   10,986      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and purchases from ESPP 2,427 $ 1 2,426    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and purchases from ESPP (In Shares)   1,094      
Stock-based compensation 9,355   9,355    
Balance at Dec. 31, 2017 38,940 $ 115 760,225 (97) (721,303)
Balance (in shares) at Dec. 31, 2017   115,555      
Net loss (57,564)       (57,564)
Other comprehensive loss (184)     (184)  
Issuance of common stock from public offering, net of offering costs 85,085 $ 18 85,067    
Issuance of common stock from public offering, net of offering costs (in shares)   18,202      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and purchases from ESPP 7,848 $ 3 7,845    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and purchases from ESPP (In Shares)   3,096      
Stock-based compensation 10,394   10,394    
Balance at Dec. 31, 2018 $ 84,519 $ 136 $ 863,531 $ (281) $ (778,867)
Balance (in shares) at Dec. 31, 2018   136,853