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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
shares in Thousands, $ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Other Comprehensive Income (Loss)
Accumulated Deficit
Balance at Dec. 31, 2017 $ 38,940 $ 115 $ 760,225 $ (97) $ (721,303)
Balance (in shares) at Dec. 31, 2017   115,555      
Issuance of common stock from public offering, net of offering costs 57,262 $ 15 57,247    
Issuance of common stock from public offering, net of offering costs (in shares)   14,039      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 1,295 $ 1 1,294    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   951      
Stock-based compensation 2,315   2,315    
Other comprehensive income (329)     (329)  
Net loss (13,885)       (13,885)
Balance at Mar. 31, 2018 85,598 $ 131 821,081 (426) (735,188)
Balance (in shares) at Mar. 31, 2018   130,545      
Balance at Dec. 31, 2017 38,940 $ 115 760,225 (97) (721,303)
Balance (in shares) at Dec. 31, 2017   115,555      
Net loss (41,359)        
Balance at Sep. 30, 2018 91,801 $ 136 854,653 (326) (762,662)
Balance (in shares) at Sep. 30, 2018   135,593      
Balance at Mar. 31, 2018 85,598 $ 131 821,081 (426) (735,188)
Balance (in shares) at Mar. 31, 2018   130,545      
Issuance of common stock from public offering, net of offering costs 11,910 $ 2 11,908    
Issuance of common stock from public offering, net of offering costs (in shares)   1,780      
Issuance of common stock from exercise of stock options and vesting of restricted stock units 3,616   3,616    
Issuance of common stock from exercise of stock options and vesting of restricted stock units (in shares)   1,137      
Stock-based compensation 2,593   2,593    
Other comprehensive income 9     9  
Net loss (13,282)       (13,282)
Balance at Jun. 30, 2018 90,444 $ 133 839,198 (417) (748,470)
Balance (in shares) at Jun. 30, 2018   133,462      
Issuance of common stock from public offering, net of offering costs 10,319 $ 1 10,318    
Issuance of common stock from public offering, net of offering costs (in shares)   1,411      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 2,468 $ 2 2,466    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   720      
Stock-based compensation 2,671   2,671    
Other comprehensive income 91     91  
Net loss (14,192)       (14,192)
Balance at Sep. 30, 2018 91,801 $ 136 854,653 (326) (762,662)
Balance (in shares) at Sep. 30, 2018   135,593      
Balance at Dec. 31, 2018 84,519 $ 136 863,531 (281) (778,867)
Balance (in shares) at Dec. 31, 2018   136,853      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 684 $ 2 682    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   965      
Stock-based compensation 2,878   2,878    
Other comprehensive income 211     211  
Net loss (18,792)       (18,792)
Balance at Mar. 31, 2019 69,500 $ 138 867,091 (70) (797,659)
Balance (in shares) at Mar. 31, 2019   137,818      
Balance at Dec. 31, 2018 84,519 $ 136 863,531 (281) (778,867)
Balance (in shares) at Dec. 31, 2018   136,853      
Net loss (54,321)        
Balance at Sep. 30, 2019 60,808 $ 142 893,715 139 (833,188)
Balance (in shares) at Sep. 30, 2019   142,010      
Balance at Mar. 31, 2019 69,500 $ 138 867,091 (70) (797,659)
Balance (in shares) at Mar. 31, 2019   137,818      
Issuance of common stock from public offering, net of offering costs 9,836 $ 2 9,834    
Issuance of common stock from public offering, net of offering costs (in shares)   1,978      
Issuance of common stock from exercise of stock options and vesting of restricted stock units 558   558    
Issuance of common stock from exercise of stock options and vesting of restricted stock units (in shares)   265      
Stock-based compensation 3,139   3,139    
Other comprehensive income 161     161  
Net loss (17,562)       (17,562)
Balance at Jun. 30, 2019 65,632 $ 140 880,622 91 (815,221)
Balance (in shares) at Jun. 30, 2019   140,061      
Issuance of common stock from public offering, net of offering costs 7,914 $ 1 7,913    
Issuance of common stock from public offering, net of offering costs (in shares)   1,471      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 1,586 $ 1 1,585    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   478      
Stock-based compensation 3,595   3,595    
Other comprehensive income 48     48  
Net loss (17,967)       (17,967)
Balance at Sep. 30, 2019 $ 60,808 $ 142 $ 893,715 $ 139 $ (833,188)
Balance (in shares) at Sep. 30, 2019   142,010