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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
shares in Thousands, $ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Other Comprehensive Income (Loss)
Accumulated Deficit
Balance at Dec. 31, 2018 $ 84,519 $ 136 $ 863,531 $ (281) $ (778,867)
Balance (in shares) at Dec. 31, 2018   136,853      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 684 $ 2 682    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   965      
Stock-based compensation 2,878   2,878    
Other comprehensive income (loss) 211     211  
Net loss (18,792)       (18,792)
Balance at Mar. 31, 2019 69,500 $ 138 867,091 (70) (797,659)
Balance (in shares) at Mar. 31, 2019   137,818      
Balance at Dec. 31, 2018 84,519 $ 136 863,531 (281) (778,867)
Balance (in shares) at Dec. 31, 2018   136,853      
Net loss (54,321)        
Balance at Sep. 30, 2019 60,808 $ 142 893,715 139 (833,188)
Balance (in shares) at Sep. 30, 2019   142,010      
Balance at Mar. 31, 2019 69,500 $ 138 867,091 (70) (797,659)
Balance (in shares) at Mar. 31, 2019   137,818      
Issuance of common stock from public offerings, net of offering costs 9,836 $ 2 9,834    
Issuance of common stock from public offering, net of offering costs (in shares)   1,978      
Issuance of common stock from exercise of stock options and vesting of restricted stock units 558   558    
Issuance of common stock from exercise of stock options and vesting of restricted stock units (in shares)   265      
Stock-based compensation 3,139   3,139    
Other comprehensive income (loss) 161     161  
Net loss (17,562)       (17,562)
Balance at Jun. 30, 2019 65,632 $ 140 880,622 91 (815,221)
Balance (in shares) at Jun. 30, 2019   140,061      
Issuance of common stock from public offerings, net of offering costs 7,914 $ 1 7,913    
Issuance of common stock from public offering, net of offering costs (in shares)   1,471      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 1,586 $ 1 1,585    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   478      
Stock-based compensation 3,595   3,595    
Other comprehensive income (loss) 48     48  
Net loss (17,967)       (17,967)
Balance at Sep. 30, 2019 60,808 $ 142 893,715 139 (833,188)
Balance (in shares) at Sep. 30, 2019   142,010      
Balance at Dec. 31, 2019 57,052 $ 144 906,905 114 (850,111)
Balance (in shares) at Dec. 31, 2019   144,291      
Issuance of common stock from public offerings, net of offering costs 67,112 $ 18 67,094    
Issuance of common stock from public offering, net of offering costs (in shares)   17,867      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 1,193 $ 2 1,191    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   1,803      
Stock-based compensation 3,754   3,754    
Other comprehensive income (loss) (237)     (237)  
Net loss (16,466)       (16,466)
Balance at Mar. 31, 2020 112,408 $ 164 978,944 (123) (866,577)
Balance (in shares) at Mar. 31, 2020   163,961      
Balance at Dec. 31, 2019 57,052 $ 144 906,905 114 (850,111)
Balance (in shares) at Dec. 31, 2019   144,291      
Net loss (45,476)        
Balance at Sep. 30, 2020 109,235 $ 167 1,003,748 907 (895,587)
Balance (in shares) at Sep. 30, 2020   167,074      
Balance at Mar. 31, 2020 112,408 $ 164 978,944 (123) (866,577)
Balance (in shares) at Mar. 31, 2020   163,961      
Issuance of common stock from public offerings, net of offering costs 9,429 $ 1 9,428    
Issuance of common stock from public offering, net of offering costs (in shares)   1,471      
Issuance of common stock from exercise of stock options and vesting of restricted stock units 1,605 $ 1 1,604    
Issuance of common stock from exercise of stock options and vesting of restricted stock units (in shares)   580      
Stock-based compensation 4,706   4,706    
Other comprehensive income (loss) 1,248     1,248  
Net loss (14,871)       (14,871)
Balance at Jun. 30, 2020 114,525 $ 166 994,682 1,125 (881,448)
Balance (in shares) at Jun. 30, 2020   166,012      
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases 4,221 $ 1 4,220    
Issuance of common stock from exercise of stock options, vesting of restricted stock units, and ESPP purchases (in shares)   1,062      
Stock-based compensation 4,846   4,846    
Other comprehensive income (loss) (218)     (218)  
Net loss (14,139)       (14,139)
Balance at Sep. 30, 2020 $ 109,235 $ 167 $ 1,003,748 $ 907 $ (895,587)
Balance (in shares) at Sep. 30, 2020   167,074