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Condensed Consolidated Statements of Cash Flows - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2020
Mar. 31, 2019
Cash flows from operating activities:    
Net loss $ (3,598) $ (13,112)
Adjustments to reconcile net loss to net cash used in operating activities    
Stock-based compensation expense 710 2,076
Depreciation expense 76 51
Loss on disposal of fixed assets 2  
Unrealized foreign currency transaction losses (gains) 29 (14)
Amortization of discount on marketable debt securities (52) (283)
Changes in operating assets and liabilities:    
Prepaid expenses and other current assets 418 469
Tax incentive receivable   (94)
Accounts payable (310) (445)
Accrued restructuring costs (1,728)  
Accrued expenses (163) (1,119)
Net cash used in operating activities (4,616) (12,471)
Cash flows from investing activities:    
Proceeds from sales and maturities of marketable debt securities 20,532 34,275
Purchases of marketable debt securities (2,339) (33,466)
Purchases of property and equipment   (2)
Net cash provided by investing activities 18,193 807
Cash flows from financing activities:    
Repayment of notes payable (1,818)  
Proceeds from exercise of common stock options and employee stock purchase plan 10 95
Net cash (used in) provided by financing activities (1,808) 95
Net increase (decrease) in cash, cash equivalents and restricted cash 11,769 (11,569)
Cash, cash equivalents and restricted cash at beginning of period 28,550 49,331
Cash, cash equivalents and restricted cash at end of period 40,319 37,762
Supplemental disclosure of non-cash investing and financing activities:    
Property and equipment included in accounts payable   11
Supplemental disclosure of cash flow information:    
Cash paid for interest $ 212 $ 339