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Consolidated Statements of Stockholders' Equity - USD ($) | Common Stock [Member] | Additional Paid-in Capital [Member] | Accumulated Deficit [Member] | Total
--- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2016 | $ 403,000 | $ 431,168,000 | $ (332,473,000) | $ 99,098,000
Beginning balance, shares at Dec. 31, 2016 | 40,254,457 |  |  |
Stock option-based compensation expense |  | 4,644,000 |  | 4,644,000
Restricted stock-based compensation expense |  | 1,272,000 |  | 1,272,000
Issuance of common stock in connection with public offering, net of offering costs | $ 58,000 | 86,116,000 |  | 86,174,000
Issuance of common stock in connection with public offering, net of offering costs, shares | 5,750,000 |  |  |
Issuance of common stock in connection with the exercise of stock options |  | 344,000 |  | 344,000
Issuance of common stock in connection with the exercise of stock options, shares | 81,719 |  |  |
Issuance of common stock in connection with the exercise of $8.00 warrants | $ 6,000 | 4,994,000 |  | 5,000,000
Issuance of common stock in connection with the exercise of $8.00 warrants, shares | 625,000 |  |  |
Issuance of common stock in connection with the exercise of $5.00 warrants | $ 2,000 | 883,000 |  | 885,000
Issuance of common stock in connection with the exercise of $5.00 warrants, shares | 176,932 |  |  |
Net loss |  |  | (27,319,000) | (27,319,000)
Ending balance at Dec. 31, 2017 | $ 469,000 | 529,421,000 | (359,792,000) | 170,098,000
Ending balance, shares at Dec. 31, 2017 | 46,888,108 |  |  |
Cumulative effect adjustment of ASC 606 on January 1, 2018 |  |  | 6,275,000 | 6,275,000
Stock option-based compensation expense |  | 8,178,000 |  | 8,178,000
Restricted stock-based compensation expense |  | 688,000 |  | 688,000
Issuance of common stock in connection with the exercise of stock options | $ 4,000 | 2,240,000 |  | 2,244,000
Issuance of common stock in connection with the exercise of stock options, shares | 360,853 |  |  |
Issuance of common stock in connection with the exercise of $5.00 warrants | $ 6,000 | 3,227,000 |  | 3,233,000
Issuance of common stock in connection with the exercise of $5.00 warrants, shares | 646,763 |  |  |
Issuance of common stock in connection with cashless warrant exercises |  |  |  |
Issuance of common stock in connection with cashless warrant exercises, shares | 48,762 |  |  |
Net loss |  |  | (56,671,000) | (56,671,000)
Ending balance at Dec. 31, 2018 | $ 479,000 | 543,754,000 | (410,188,000) | 134,045,000
Ending balance, shares at Dec. 31, 2018 | 47,944,486 |  |  |
Stock option-based compensation expense |  | 7,338,000 |  | 7,338,000
Restricted stock-based compensation expense |  | 899,000 |  | 899,000
Issuance of common stock and pre-funded warrants in connection with public offering, net of offering costs | $ 324,000 | 95,648,000 |  | 95,972,000
Issuance of common stock and pre-funded warrants in connection with public offering, net of offering costs, shares | 32,382,945 |  |  |
Issuance of common stock under open market sale agreement | $ 31,000 | 16,930,000 |  | 16,961,000
Issuance of common stock under open market sale agreement, shares | 3,086,950 |  |  |
Issuance of common stock in connection with the exercise of stock options | $ 3,000 | 966,000 |  | 969,000
Issuance of common stock in connection with the exercise of stock options, shares | 303,129 |  |  |
Issuance of common stock in connection with restricted share awards | $ 4,000 | (4,000) |  |
Issuance of common stock in connection with restricted share awards, shares | 354,625 |  |  |
Shares returned in connection with arbitration ruling on licensing agreement | $ (5,000) | (1,467,000) |  | (1,472,000)
Shares returned in connection with arbitration ruling on licensing agreement, shares | (450,000) |  |  |
Net loss |  |  | (76,282,000) | (76,282,000)
Ending balance at Dec. 31, 2019 | $ 836,000 | $ 664,064,000 | $ (486,470,000) | $ 178,430,000
Ending balance, shares at Dec. 31, 2019 | 83,622,135 |  |  |

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