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Revenue Arrangements - Narrative (Details) |  |  |  |  | 3 Months Ended |
--- | --- | --- | --- | --- | --- | ---
Jul. 28, 2020 USD ($) | Jun. 13, 2020 USD ($) | Jun. 11, 2020 USD ($) obligation | Dec. 17, 2019 USD ($) | Mar. 31, 2021 USD ($) | Mar. 31, 2020 USD ($) | Dec. 31, 2020 USD ($)
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |
Short-term contract liability |  |  |  |  | $ 75,764,000 |  | $ 72,050,000
Long-term contract liability |  |  |  |  | 32,301,000 |  | 38,746,000
Contract assets amortization (less than) |  |  |  |  | 100,000 |  |
Accounts receivable |  |  |  |  | 8,342,000 |  | 7,224,000
Revenue recognized |  |  |  |  | 2,730,000 |  |
Remaining performance obligation |  |  |  |  | 108,100,000 |  |
SOBI Purchase Agreement |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |
Sale common stock |  |  | $ 25,000,000.0 |  |  |  |
Redemption premium | $ 14,500,000 |  |  |  |  |  |
Short-term contract liability |  |  |  |  | 72,000,000.0 |  | 68,300,000
Long-term contract liability |  |  |  |  | 17,800,000 |  | 24,200,000
Contract assets |  |  |  |  | 1,300,000 |  |
Accounts receivable |  |  |  |  | 8,300,000 |  | 6,900,000
Revenue recognized |  |  |  |  | 11,100,000 |  |
SOBI Purchase Agreement | Prepaid Expenses and Other Current Assets |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |
Contract assets |  |  |  |  | 1,000,000.0 |  |
SOBI Purchase Agreement | Other Current Assets |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |
Contract assets |  |  |  |  | 300,000 |  |
Collaborative Arrangement |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |
Upfront cash payment |  |  | 75,000,000.0 |  |  |  |
Sales milestone payments |  |  | $ 630,000,000.0 |  |  |  |
License and option agreement, written day notice period before cancellation |  |  | 180 days |  |  |  |
Period after first commercial sale when the Company is eligible to receive royalties |  |  | 10 years |  |  |  |
Number of obligations | obligation |  |  | 3 |  |  |  |
Development milestone |  |  | $ 5,000,000.0 |  |  |  |
Sarepta Therapeutics, Inc. |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |
Upfront cash payment |  | $ 2,000,000.0 |  |  |  |  |
License and option agreement, written day notice period before cancellation |  | 30 days |  |  |  |  |
Short-term contract liability |  |  |  |  | 2,000,000.0 |  | 2,000,000.0
Revenue recognized |  |  |  |  | 0 | $ 0 |
Option term |  | 24 months |  |  |  |  |
License Agreement For Pompe Disease |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |
Upfront cash payment |  |  |  | $ 7,000,000.0 |  |  |
Sales milestone payments |  |  |  | $ 237,000,000.0 |  |  |
Period after first commercial sale when the Company is eligible to receive royalties |  |  |  | 10 years |  |  |
Short-term contract liability |  |  |  |  | 1,700,000 |  | 1,700,000
Long-term contract liability |  |  |  |  | 5,300,000 |  | $ 5,300,000
Revenue recognized |  |  |  |  | $ 0 | $ 0 |
Initial up-front cash payment |  |  |  | $ 2,000,000.0 |  |  |
Initial upfront execution fee payment |  |  |  | $ 5,000,000.0 |  |  |

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