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Consolidated Statements of Operations and Comprehensive Loss - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2021
Mar. 31, 2020
Statement of Comprehensive Income [Abstract]    
Grant and collaboration revenue $ 11,050 $ 0
Operating expenses:    
Research and development 13,004 14,724
General and administrative 5,204 4,098
Total operating expenses 18,208 18,822
Loss from operations (7,158) (18,822)
Investment income 12 240
Foreign currency transaction gain, net 7 82
Interest expense (711) (273)
Change in fair value of warrant liabilities (16,747) (846)
Other income (expense), net 0 (1)
Net loss (24,597) (19,620)
Other comprehensive loss:    
Foreign currency translation adjustment (6) (60)
Unrealized (losses) on marketable securities (1) 0
Total comprehensive loss $ (24,604) $ (19,680)
Net loss per share:    
Basic and diluted (in dollars per share) $ (0.22) $ (0.21)
Weighted average common shares outstanding:    
Basic and diluted (in shares) 110,742,150 94,723,513