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Revenue Arrangements - Narrative (Details) - USD ($) |  | 1 Months Ended | 3 Months Ended | 9 Months Ended |  |
--- | --- | --- | --- | --- | --- | ---
Jun. 28, 2024 | Jan. 31, 2023 | Oct. 31, 2021 | Jun. 30, 2020 | Sep. 30, 2024 | Sep. 30, 2023 | Jun. 30, 2023 | Sep. 30, 2024 | Sep. 30, 2023 | Jun. 30, 2024 | Dec. 31, 2023
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Deferred revenue related to agreement |  |  |  |  | $ 0 |  |  | $ 0 |  |  | $ 5,849,000
Deferred revenue |  |  |  |  | 0 |  |  | 0 |  |  | 2,311,000
Deferred revenue, net of current portion |  |  |  |  | 0 |  |  | 0 |  |  | 3,538,000
Accounts receivable |  |  |  |  | 4,829,000 |  |  | 4,829,000 |  |  | 5,870,000
Revenue recognized |  |  |  |  |  |  |  | (5,849,000) |  |  |
Collaboration and license revenue |  |  |  |  | 0 | $ 6,551,000 |  | 39,111,000 | $ 17,738,000 |  |
Remaining performance obligation |  |  |  |  | 0 |  |  | 0 |  |  |
Grant revenue |  |  |  |  | 387,000 | 0 |  | 561,000 | 0 |  |
National Institute of Neurological Disorders and Stroke of the National Institutes of Health Funding |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Government assistance, award amount |  |  |  |  |  |  |  |  |  | $ 1,500,000 |
Government assistance, potential additional award amount |  |  |  |  |  |  |  |  |  | $ 1,500,000 |
Government assistance, asset, current |  |  |  |  | 600,000 |  |  | 600,000 |  |  |
Grant revenue |  |  |  |  | 400,000 |  |  | 600,000 |  |  |
SOBI Purchase Agreement |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Accounts receivable |  |  |  |  | 4,200,000 |  |  | 4,200,000 |  |  | 4,600,000
Collaboration and license revenue |  |  |  |  | 0 | 5,000,000 |  | 32,800,000 | 13,700,000 |  |
SOBI Purchase Agreement | Billed Revenues |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Accounts receivable |  |  |  |  | 1,000,000.0 |  |  | 1,000,000.0 |  |  | 3,000,000.0
Takeda Agreement |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  | 0 |  |  | 0 |  |  | 0
Upfront cash payment |  |  | $ 3,000,000 |  |  |  |  |  |  |  |
Future additional payments, expected |  |  | $ 1,124,000,000 |  |  |  |  |  |  |  |
Milestone payments |  |  |  |  |  |  | $ 500,000 |  |  |  |
Customer, liability, revenue recognized, including opening balance |  |  |  |  | 0 | 0 |  | 0 | 600,000 |  |
Collaborative Arrangement | SOBI Purchase Agreement |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Upfront cash payment |  |  |  | $ 75,000,000 |  |  |  |  |  |  |
Sales milestone payments |  |  |  | $ 630,000,000 |  |  |  |  |  |  |
Revenue recognized | $ 30,000,000 |  |  |  |  |  |  | 30,000,000 |  |  |
Astellas Gene Therapies |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Upfront payment |  | $ 10,000,000 |  |  |  |  |  |  |  |  |
Development and commercial milestones plus royalties |  | $ 340,000,000 |  |  |  |  |  |  |  |  |
Deferred revenue related to agreement |  |  |  |  | 0 |  |  | 0 |  |  |
Deferred revenue |  |  |  |  |  |  |  |  |  |  | 2,300,000
Deferred revenue, net of current portion |  |  |  |  |  |  |  |  |  |  | 3,500,000
Accounts receivable |  |  |  |  | 100,000 |  |  | 100,000 |  |  | $ 300,000
Revenue recognized |  |  |  |  | 0 | 1,500,000 |  | 6,300,000 | 2,900,000 |  |
Revenue, remaining performance obligation recognized |  |  |  |  |  |  |  | 3,200,000 |  |  |
Sarepta Therapeutics, Inc. |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |
Revenue recognized |  |  |  |  | $ 0 | $ 0 |  | $ 0 | $ 500,000 |  |
Option term |  |  |  | 24 months |  |  |  |  |  |  |

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