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Revenue Arrangements - Narrative (Details) |  |  |  |  |  |  | 1 Months Ended | 3 Months Ended | 12 Months Ended |
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Jun. 28, 2024 USD ($) | Jun. 29, 2022 USD ($) | Jun. 10, 2022 | Oct. 01, 2021 USD ($) | Jul. 28, 2020 USD ($) | Jun. 11, 2020 USD ($) obligation | Jan. 31, 2023 USD ($) | Jun. 30, 2020 USD ($) | Jun. 30, 2021 USD ($) | Dec. 31, 2024 USD ($) | Dec. 31, 2023 USD ($) | Dec. 31, 2022 USD ($) | Jun. 30, 2024 USD ($)
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Short-term contract liability |  |  |  |  |  |  |  |  |  | $ 0 | $ 2,311,000 |  |
Long-term contract liability |  |  |  |  |  |  |  |  |  | 0 | 3,538,000 |  |
Accounts receivable |  |  |  |  |  |  |  |  |  | 872,000 | 5,870,000 |  |
Revenue recognized |  |  |  |  |  |  |  |  |  | 5,849,000 |  |  |
Collaboration and license revenue |  |  |  |  |  |  |  |  |  | 38,275,000 | 26,004,000 |  |
Remaining performance obligation |  |  |  |  |  |  |  |  |  | $ 0 |  |  |
Government Assistance Current Statement Of Financial Position Extensible Enumeration Not Disclosed Flag |  |  |  |  |  |  |  |  |  | receivable |  |  |
Grant revenue |  |  |  |  |  |  |  |  |  | $ 638,000 | 0 |  |
National Institute of Neurological Disorders and Stroke of the National Institutes of Health Funding |  |  |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Government assistance, award amount |  |  |  |  |  |  |  |  |  |  |  |  | $ 1,500,000
Government assistance, potential additional award amount |  |  |  |  |  |  |  |  |  |  |  |  | $ 1,500,000
Government assistance, asset, current |  |  |  |  |  |  |  |  |  | 600,000 |  |  |
Grant revenue |  |  |  |  |  |  |  |  |  | 600,000 |  |  |
SOBI Purchase Agreement |  |  |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Sale common stock |  |  |  |  |  | $ 25,000,000 |  |  |  |  |  |  |
Redemption premium |  |  |  |  | $ 14,500,000 |  |  |  |  |  |  |  |
Astellas Gene Therapies |  |  |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Collaboration and license agreements upfront payment |  |  |  |  |  |  | $ 10,000,000 |  |  |  |  |  |
Development and commercial milestones plus royalties |  |  |  |  |  |  | $ 340,000,000 |  |  |  |  |  |
Reimbursement percentage, budgeted costs incurred to complete the development |  |  |  |  |  |  | 25.00% |  |  |  |  |  |
License and option agreement, development cost reimbursements |  |  |  |  |  |  | $ 5,500,000 |  |  |  |  |  |
License and option agreement, written notice period before cancellation |  |  |  |  |  |  | 90 days |  |  |  |  |  |
Short-term contract liability |  |  |  |  |  |  |  |  |  | 0 | 2,300,000 |  |
Long-term contract liability |  |  |  |  |  |  |  |  |  |  | 3,500,000 |  |
Accounts receivable |  |  |  |  |  |  |  |  |  | 100,000 | 300,000 |  |
Customer, liability, revenue recognized, including opening balance |  |  |  |  |  |  |  |  |  | 6,300,000 |  |  |
Revenue, remaining performance obligation recognized |  |  |  |  |  |  |  |  |  | 3,200,000 |  |  |
Revenue recognized |  |  |  |  |  |  |  |  |  |  | 5,500,000 |  |
Sarepta Therapeutics, Inc. |  |  |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
License and option agreement, written notice period before cancellation |  |  |  |  |  |  |  | 30 days |  |  |  |  |
Short-term contract liability |  |  |  |  |  |  |  |  |  | 0 | 0 |  |
Upfront cash payment |  |  |  |  |  |  |  | $ 2,000,000.0 |  |  |  |  |
Sales milestone payments |  |  |  |  |  |  |  |  | $ 3,000,000 |  |  |  |
Collaboration and license revenue |  |  |  |  |  |  |  |  |  | 0 | 500,000 |  |
Option term |  |  |  |  |  |  |  | 24 months |  |  |  |  |
Extension term |  |  | 9 months |  |  |  |  |  |  |  |  |  |
Milestone receivable for certain dystrophies |  |  |  |  |  |  |  |  |  |  |  | $ 2,000,000 |
Milestone receivable |  |  |  |  |  |  |  |  |  |  |  | $ 4,000,000 |
Takeda Agreement |  |  |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
License and option agreement, written notice period before cancellation |  |  |  | 90 days |  |  |  |  |  |  |  |  |
Short-term contract liability |  |  |  |  |  |  |  |  |  | 0 | 0 |  |
Contract with customer, asset, reclassified to receivable |  |  |  |  |  |  |  |  |  |  | 500,000 |  |
Customer, liability, revenue recognized, including opening balance |  |  |  |  |  |  |  |  |  | 0 | 600,000 |  |
Upfront cash payment |  |  |  | $ 3,000,000 |  |  |  |  |  |  |  |  |
Future additional payments, expected |  |  |  | $ 1,124,000,000 |  |  |  |  |  |  |  |  |
Collaborative Arrangement | Swedish Orphan Biovitrum AB (SOBI) |  |  |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
License and option agreement, written notice period before cancellation |  |  |  |  |  | 180 days |  |  |  |  |  |  |
Accounts receivable |  |  |  |  |  |  |  |  |  | 100,000 | 4,600,000 |  |
Revenue, remaining performance obligation recognized |  |  |  |  |  |  |  |  |  | 1,900,000 | 1,100,000 |  |
Upfront cash payment |  |  |  |  |  | $ 75,000,000 |  |  |  |  |  |  |
Sales milestone payments |  |  |  |  |  | $ 630,000,000 |  |  |  |  |  |  |
Period after first commercial sale when the Company is eligible to receive royalties |  |  |  |  |  | 10 years |  |  |  |  |  |  |
Number of obligations | obligation |  |  |  |  |  | 3 |  |  |  |  |  |  |
Development milestone |  |  |  |  |  | $ 5,000,000 |  |  |  |  |  |  |
Collaborative arrangement, payment obligation |  | $ 10,000,000 |  |  |  |  |  |  |  |  |  |  |
Revenue recognized | $ 30,000,000 |  |  |  |  |  |  |  |  | 30,000,000 |  |  |
Collaboration and license revenue |  |  |  |  |  |  |  |  |  | 31,900,000 | 19,400,000 |  |
Collaborative Arrangement | Billed Revenues | Swedish Orphan Biovitrum AB (SOBI) |  |  |  |  |  |  |  |  |  |  |  |  |
Disaggregation of Revenue [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Accounts receivable |  |  |  |  |  |  |  |  |  | $ 0 | $ 3,000,000.0 |  |

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