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Condensed Consolidated Statements of Stockholders' Equity (Unaudited) - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Other Comprehensive (Loss) Income | Accumulated Deficit
--- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2017 | $ 131,136 | $ 646 | $ 913,955 | $ (10) | $ (783,455)
Balance (in shares) at Dec. 31, 2017 |  | 64,609,000 |  |  |
Cumulative effect of adoption of new accounting standard at Dec. 31, 2017 | 1,573 |  |  |  | 1,573
Issuance of common stock in public offerings, net | (205) |  | (205) |  |
Conversion benefit included in Convertible Notes issued | 96 |  | 96 |  |
Issuance of common stock on exercise of stock options | 4,202 | $ 4 | 4,198 |  |
Issuance of common stock on exercise of stock options (in shares) |  | 435,000 |  |  |
Stock-based compensation expense | 7,701 |  | 7,701 |  |
Net loss | (52,265) |  |  |  | (52,265)
Net unrealized gain (loss) on short-term investments | (32) |  |  | (32) |
Comprehensive loss | (52,297) |  |  |  |
Balance at Mar. 31, 2018 | 92,206 | $ 650 | 925,745 | (42) | (834,147)
Balance (in shares) at Mar. 31, 2018 |  | 65,044,000 |  |  |
Cumulative effect of adoption of new accounting standard at Mar. 31, 2018 | 1 |  |  |  | 1
Balance at Dec. 31, 2017 | 131,136 | $ 646 | 913,955 | (10) | (783,455)
Balance (in shares) at Dec. 31, 2017 |  | 64,609,000 |  |  |
Cumulative effect of adoption of new accounting standard at Dec. 31, 2017 | 1,573 |  |  |  | 1,573
Net loss | (129,281) |  |  |  |
Net unrealized gain (loss) on short-term investments | (32) |  |  |  |
Comprehensive loss | (129,313) |  |  |  |
Balance at Sep. 30, 2018 | 406,808 | $ 780 | 1,317,232 | (42) | (911,162)
Balance (in shares) at Sep. 30, 2018 |  | 77,953,000 |  |  |
Balance at Mar. 31, 2018 | 92,206 | $ 650 | 925,745 | (42) | (834,147)
Balance (in shares) at Mar. 31, 2018 |  | 65,044,000 |  |  |
Cumulative effect of adoption of new accounting standard at Mar. 31, 2018 | 1 |  |  |  | 1
Issuance of common stock in public offerings, net | 363,332 | $ 120 | 363,212 |  |
Issuance of common stock in public offerings, net (in shares) |  | 11,963,000 |  |  |
Conversion benefit included in Convertible Notes issued | 97 |  | 97 |  |
Issuance of common stock on exercise of stock options | 6,439 | $ 5 | 6,434 |  |
Issuance of common stock on exercise of stock options (in shares) |  | 504,000 |  |  |
Issuance of common stock on under the employee stock purchase plan | 648 |  | 648 |  |
Issuance of common stock on under the Employee Stock Purchase Plan (in shares) |  | 49,000 |  |  |
Stock-based compensation expense | 7,821 |  | 7,821 |  |
Net loss | (38,670) |  |  |  | (38,670)
Net unrealized gain (loss) on short-term investments | 17 |  |  | 17 |
Comprehensive loss | (38,653) |  |  |  |
Balance at Jun. 30, 2018 | 431,891 | $ 775 | 1,303,957 | (25) | (872,816)
Balance (in shares) at Jun. 30, 2018 |  | 77,560,000 |  |  |
Issuance of common stock in public offerings, net | 1 |  | 1 |  |
Conversion benefit included in Convertible Notes issued | 99 |  | 99 |  |
Issuance of common stock on exercise of stock options | 5,130 | $ 5 | 5,125 |  |
Issuance of common stock on exercise of stock options (in shares) |  | 393,000 |  |  |
Stock-based compensation expense | 8,050 |  | 8,050 |  |
Net loss | (38,346) |  |  |  | (38,346)
Net unrealized gain (loss) on short-term investments | (17) |  |  | (17) |
Comprehensive loss | (38,363) |  |  |  |
Balance at Sep. 30, 2018 | 406,808 | $ 780 | 1,317,232 | (42) | (911,162)
Balance (in shares) at Sep. 30, 2018 |  | 77,953,000 |  |  |
Balance at Dec. 31, 2018 | 370,160 | $ 782 | 1,330,186 | (87) | (960,721)
Balance (in shares) at Dec. 31, 2018 |  | 78,174,000 |  |  |
Conversion benefit included in Convertible Notes issued | 102 |  | 102 |  |
Issuance of common stock on exercise of stock options | 6,539 | $ 7 | 6,532 |  |
Issuance of common stock on exercise of stock options (in shares) |  | 732,000 |  |  |
Stock-based compensation expense | 17,902 |  | 17,902 |  |
Net loss | (63,012) |  |  |  | (63,012)
Net unrealized gain (loss) on short-term investments | 123 |  |  | 123 |
Comprehensive loss | (62,889) |  |  |  |
Balance at Mar. 31, 2019 | 331,814 | $ 789 | 1,354,722 | 36 | (1,023,733)
Balance (in shares) at Mar. 31, 2019 |  | 78,906,000 |  |  |
Balance at Dec. 31, 2018 | $ 370,160 | $ 782 | 1,330,186 | (87) | (960,721)
Balance (in shares) at Dec. 31, 2018 |  | 78,174,000 |  |  |
Issuance of common stock on exercise of stock options (in shares) | 1,793,000 |  |  |  |
Net loss | $ (146,829) |  |  |  |
Net unrealized gain (loss) on short-term investments | 190 |  |  |  |
Comprehensive loss | (146,639) |  |  |  |
Balance at Sep. 30, 2019 | 285,442 | $ 800 | 1,392,089 | 103 | (1,107,550)
Balance (in shares) at Sep. 30, 2019 |  | 80,030,000 |  |  |
Balance at Mar. 31, 2019 | 331,814 | $ 789 | 1,354,722 | 36 | (1,023,733)
Balance (in shares) at Mar. 31, 2019 |  | 78,906,000 |  |  |
Conversion benefit included in Convertible Notes issued | 103 |  | 103 |  |
Issuance of common stock on exercise of stock options | 9,676 | $ 8 | 9,668 |  |
Issuance of common stock on exercise of stock options (in shares) |  | 810,000 |  |  |
Issuance of common stock on under the employee stock purchase plan | 1,170 | $ 1 | 1,169 |  |
Issuance of common stock on under the Employee Stock Purchase Plan (in shares) |  | 63,000 |  |  |
Stock-based compensation expense | 12,706 |  | 12,706 |  |
Net loss | (50,222) |  |  |  | (50,222)
Net unrealized gain (loss) on short-term investments | 112 |  |  | 112 |
Comprehensive loss | (50,110) |  |  |  |
Balance at Jun. 30, 2019 | 305,359 | $ 798 | 1,378,368 | 148 | (1,073,955)
Balance (in shares) at Jun. 30, 2019 |  | 79,779,000 |  |  |
Issuance of common stock in public offerings, net | (110) |  | (110) |  |
Conversion benefit included in Convertible Notes issued | 105 |  | 105 |  |
Issuance of common stock on exercise of stock options | 4,024 | $ 2 | 4,022 |  |
Issuance of common stock on exercise of stock options (in shares) |  | 251,000 |  |  |
Stock-based compensation expense | 9,704 |  | 9,704 |  |
Net loss | (33,595) |  |  |  | (33,595)
Net unrealized gain (loss) on short-term investments | (45) |  |  | (45) |
Comprehensive loss | (33,640) |  |  |  |
Balance at Sep. 30, 2019 | $ 285,442 | $ 800 | $ 1,392,089 | $ 103 | $ (1,107,550)
Balance (in shares) at Sep. 30, 2019 |  | 80,030,000 |  |  |

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