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Consolidated Statements of Stockholders' Equity (Deficit) - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Other Comprehensive (Loss) Income | Accumulated Deficit
--- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2016 | $ (21,251) | $ 394 | $ 564,343 | $ (17) | $ (585,971)
Balance (in shares) at Dec. 31, 2016 |  | 39,355,000 |  |  |
Issuance of common stock in public offerings, net | 306,279 | $ 238 | 306,041 |  |
Issuance of common stock in public offerings, net (in shares) |  | 23,822,000 |  |  |
Conversion benefit included in Convertible Notes issued | 369 |  | 369 |  |
Issuance of common stock under Employee Stock Purchase Plan | 989 | $ 1 | 988 |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  | 77,000 |  |  |
Issuance of common stock on exercise of stock options | $ 11,463 | $ 13 | 11,450 |  |
Issuance of common stock on exercise of stock options (in shares) | 1,350,605 | 1,351,000 |  |  |
Issuance of common stock on exercise of warrants (in shares) |  | 4,000 |  |  |
Issuance of warrants | $ 226 |  | 226 |  |
Stock-based compensation expense | 30,538 |  | 30,538 |  |
Net loss | (197,484) |  |  |  | (197,484)
Net unrealized gain (loss) on short-term investments | 7 |  |  | 7 |
Comprehensive loss | (197,477) |  |  |  |
Balance at Dec. 31, 2017 | 131,136 | $ 646 | 913,955 | (10) | (783,455)
Balance (in shares) at Dec. 31, 2017 |  | 64,609,000 |  |  |
Cumulative effect of adoption of new accounting standard at Dec. 31, 2017 | 1,574 |  |  |  | 1,574
Issuance of common stock in public offerings, net | 363,128 | $ 120 | 363,008 |  |
Issuance of common stock in public offerings, net (in shares) |  | 11,963,000 |  |  |
Conversion benefit included in Convertible Notes issued | 392 |  | 392 |  |
Issuance of common stock under Employee Stock Purchase Plan | 1,179 | $ 1 | 1,178 |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  | 72,000 |  |  |
Issuance of common stock on exercise of stock options | $ 18,301 | $ 15 | 18,286 |  |
Issuance of common stock on exercise of stock options (in shares) | 1,529,509 | 1,530,000 |  |  |
Stock-based compensation expense | $ 33,367 |  | 33,367 |  |
Net loss | (178,840) |  |  |  | (178,840)
Net unrealized gain (loss) on short-term investments | (77) |  |  | (77) |
Comprehensive loss | (178,917) |  |  |  |
Balance at Dec. 31, 2018 | 370,160 | $ 782 | 1,330,186 | (87) | (960,721)
Balance (in shares) at Dec. 31, 2018 |  | 78,174,000 |  |  |
Issuance of common stock in public offerings, net | 162,151 | $ 99 | 162,052 |  |
Issuance of common stock in public offerings, net (in shares) |  | 9,857,000 |  |  |
Conversion benefit included in Convertible Notes issued | 416 |  | 416 |  |
Issuance of common stock under Employee Stock Purchase Plan | 2,109 | $ 1 | 2,108 |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  | 126,000 |  |  |
Issuance of common stock on exercise of stock options | $ 22,164 | $ 20 | 22,144 |  |
Issuance of common stock on exercise of stock options (in shares) | 1,983,221 | 1,983,000 |  |  |
Issuance of common stock on exercise of warrants | $ 1 | $ 1 |  |  |
Issuance of common stock on exercise of warrants (in shares) |  | 132,000 |  |  |
Issuance of common stock on conversion of convertible notes (in shares) |  | 32,000 |  |  |
Stock-based compensation expense | 51,411 |  | 51,411 |  |
Net loss | (204,749) |  |  |  | (204,749)
Net unrealized gain (loss) on short-term investments | 172 |  |  | 172 |
Comprehensive loss | (204,577) |  |  |  |
Balance at Dec. 31, 2019 | $ 403,835 | $ 903 | $ 1,568,317 | $ 85 | $ (1,165,470)
Balance (in shares) at Dec. 31, 2019 |  | 90,304,000 |  |  |

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