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Condensed Consolidated Statements of Stockholders' Equity (Unaudited) - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Other Comprehensive (Loss) Income | Accumulated Deficit
--- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2018 | $ 370,160 | $ 782 | $ 1,330,186 | $ (87) | $ (960,721)
Balance (in shares) at Dec. 31, 2018 |  | 78,174,000 |  |  |
Conversion benefit included in Convertible Notes issued | 102 |  | 102 |  |
Issuance of common stock on exercise of stock options | 6,539 | $ 7 | 6,532 |  |
Issuance of common stock on exercise of stock options (in shares) |  | 732,000 |  |  |
Stock-based compensation expense | 17,902 |  | 17,902 |  |
Net loss | (63,012) |  |  |  | (63,012)
Net unrealized gain on short-term investments | 123 |  |  | 123 |
Comprehensive loss | (62,889) |  |  |  |
Balance at Mar. 31, 2019 | 331,814 | $ 789 | 1,354,722 | 36 | (1,023,733)
Balance (in shares) at Mar. 31, 2019 |  | 78,906,000 |  |  |
Balance at Dec. 31, 2019 | 403,835 | $ 903 | 1,568,317 | 85 | (1,165,470)
Balance (in shares) at Dec. 31, 2019 |  | 90,304,000 |  |  |
Conversion benefit included in Convertible Notes issued | 108 |  | 108 |  |
Issuance of common stock on exercise of stock options | $ 504 |  | 504 |  |
Issuance of common stock on exercise of stock options (in shares) | 70,000 | 70,000 |  |  |
Issuance of common stock on exercise of warrants | $ 3 | $ 3 |  |  |
Issuance of common stock on exercise of warrants (in shares) |  | 268,000 |  |  |
Stock-based compensation expense | 11,974 |  | 11,974 |  |
Net loss | (51,579) |  |  |  | (51,579)
Net unrealized gain on short-term investments | 623 |  |  | 623 |
Comprehensive loss | (50,956) |  |  |  |
Balance at Mar. 31, 2020 | $ 365,468 | $ 906 | $ 1,580,903 | $ 708 | $ (1,217,049)
Balance (in shares) at Mar. 31, 2020 |  | 90,642,000 |  |  |

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