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Condensed Consolidated Statements of Stockholders' Equity (Unaudited) - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-In Capital
Accumulated Other Comprehensive (Loss) Income
Accumulated Deficit
Balance at Dec. 31, 2018 $ 370,160 $ 782 $ 1,330,186 $ (87) $ (960,721)
Balance (in shares) at Dec. 31, 2018   78,174,000      
Conversion benefit included in Convertible Notes issued 102   102    
Issuance of common stock on exercise of stock options 6,539 $ 7 6,532    
Issuance of common stock on exercise of stock options (in shares)   732,000      
Stock-based compensation expense 17,902   17,902    
Net loss (63,012)       (63,012)
Net unrealized gain on short-term investments 123     123  
Comprehensive loss (62,889)        
Balance at Mar. 31, 2019 331,814 $ 789 1,354,722 36 (1,023,733)
Balance (in shares) at Mar. 31, 2019   78,906,000      
Balance at Dec. 31, 2018 370,160 $ 782 1,330,186 (87) (960,721)
Balance (in shares) at Dec. 31, 2018   78,174,000      
Net loss (113,234)        
Net unrealized gain on short-term investments 235        
Comprehensive loss (112,999)        
Balance at Jun. 30, 2019 305,359 $ 798 1,378,368 148 (1,073,955)
Balance (in shares) at Jun. 30, 2019   79,779,000      
Balance at Mar. 31, 2019 331,814 $ 789 1,354,722 36 (1,023,733)
Balance (in shares) at Mar. 31, 2019   78,906,000      
Conversion benefit included in Convertible Notes issued 103   103    
Issuance of common stock on exercise of stock options 9,676 $ 8 9,668    
Issuance of common stock on exercise of stock options (in shares)   810,000      
Issuance of common stock under the employee stock purchase plan 1,170 $ 1 1,169    
Issuance of common stock on under the Employee Stock Purchase Plan (in shares)   63,000      
Stock-based compensation expense 12,706   12,706    
Net loss (50,222)       (50,222)
Net unrealized gain on short-term investments 112     112  
Comprehensive loss (50,110)        
Balance at Jun. 30, 2019 305,359 $ 798 1,378,368 148 (1,073,955)
Balance (in shares) at Jun. 30, 2019   79,779,000      
Balance at Dec. 31, 2019 403,835 $ 903 1,568,317 85 (1,165,470)
Balance (in shares) at Dec. 31, 2019   90,304,000      
Conversion benefit included in Convertible Notes issued 108   108    
Issuance of common stock on exercise of stock options 504   504    
Issuance of common stock on exercise of stock options (in shares)   70,000      
Issuance of common stock on exercise of warrants 3 $ 3      
Issuance of common stock on exercise of warrants (in shares)   268,000      
Stock-based compensation expense 11,974   11,974    
Net loss (51,579)       (51,579)
Net unrealized gain on short-term investments 623     623  
Comprehensive loss (50,956)        
Balance at Mar. 31, 2020 365,468 $ 906 1,580,903 708 (1,217,049)
Balance (in shares) at Mar. 31, 2020   90,642,000      
Balance at Dec. 31, 2019 $ 403,835 $ 903 1,568,317 85 (1,165,470)
Balance (in shares) at Dec. 31, 2019   90,304,000      
Issuance of common stock on exercise of stock options (in shares) 124,000        
Net loss $ (106,769)        
Net unrealized gain on short-term investments 868        
Comprehensive loss (105,901)        
Balance at Jun. 30, 2020 324,058 $ 908 1,594,436 953 (1,272,239)
Balance (in shares) at Jun. 30, 2020   90,820,000      
Balance at Mar. 31, 2020 365,468 $ 906 1,580,903 708 (1,217,049)
Balance (in shares) at Mar. 31, 2020   90,642,000      
Conversion benefit included in Convertible Notes issued 109   109    
Issuance of common stock on exercise of stock options 805 $ 1 804    
Issuance of common stock on exercise of stock options (in shares)   54,000      
Issuance of common stock under the employee stock purchase plan 1,507 $ 1 1,506    
Issuance of common stock on under the Employee Stock Purchase Plan (in shares)   124,000      
Stock-based compensation expense 11,114   11,114    
Net loss (55,190)       (55,190)
Net unrealized gain on short-term investments 245     245  
Comprehensive loss (54,945)        
Balance at Jun. 30, 2020 $ 324,058 $ 908 $ 1,594,436 $ 953 $ (1,272,239)
Balance (in shares) at Jun. 30, 2020   90,820,000