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Condensed Consolidated Statements of Stockholders' Equity (Unaudited) - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-In Capital | Accumulated Other Comprehensive Income | Accumulated Deficit
--- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2019 | $ 403,835 | $ 903 | $ 1,568,317 | $ 85 | $ (1,165,470)
Balance (in shares) at Dec. 31, 2019 |  | 90,304,000 |  |  |
Conversion benefit included in Convertible Notes issued | 108 |  | 108 |  |
Issuance of common stock under equity incentive plan | 504 |  | 504 |  |
Issuance of common stock under equity incentive plan (in shares) |  | 70,000 |  |  |
Issuance of common stock on exercise of warrants | 3 | $ 3 |  |  |
Issuance of common stock on exercise of warrants (in shares) |  | 268,000 |  |  |
Stock-based compensation expense | 11,974 |  | 11,974 |  |
Net loss | (51,579) |  |  |  | (51,579)
Net unrealized gain (loss) on short-term investments | 623 |  |  | 623 |
Comprehensive loss | (50,956) |  |  |  |
Balance at Mar. 31, 2020 | 365,468 | $ 906 | 1,580,903 | 708 | (1,217,049)
Balance (in shares) at Mar. 31, 2020 |  | 90,642,000 |  |  |
Balance at Dec. 31, 2019 | 403,835 | $ 903 | 1,568,317 | 85 | (1,165,470)
Balance (in shares) at Dec. 31, 2019 |  | 90,304,000 |  |  |
Net loss | (106,769) |  |  |  |
Net unrealized gain (loss) on short-term investments | 868 |  |  |  |
Comprehensive loss | (105,901) |  |  |  |
Balance at Jun. 30, 2020 | 324,058 | $ 908 | 1,594,436 | 953 | (1,272,239)
Balance (in shares) at Jun. 30, 2020 |  | 90,820,000 |  |  |
Balance at Mar. 31, 2020 | 365,468 | $ 906 | 1,580,903 | 708 | (1,217,049)
Balance (in shares) at Mar. 31, 2020 |  | 90,642,000 |  |  |
Conversion benefit included in Convertible Notes issued | 109 |  | 109 |  |
Issuance of common stock under equity incentive plan | 805 | $ 1 | 804 |  |
Issuance of common stock under equity incentive plan (in shares) |  | 54,000 |  |  |
Issuance of common stock on under the employee stock purchase plan | 1,507 | $ 1 | 1,506 |  |
Issuance of common stock on under the employee stock purchase plan (in shares) |  | 124,000 |  |  |
Stock-based compensation expense | 11,114 |  | 11,114 |  |
Net loss | (55,190) |  |  |  | (55,190)
Net unrealized gain (loss) on short-term investments | 245 |  |  | 245 |
Comprehensive loss | (54,945) |  |  |  |
Balance at Jun. 30, 2020 | 324,058 | $ 908 | 1,594,436 | 953 | (1,272,239)
Balance (in shares) at Jun. 30, 2020 |  | 90,820,000 |  |  |
Balance at Dec. 31, 2020 | 236,492 | $ 913 | 1,628,070 | 257 | (1,392,748)
Balance (in shares) at Dec. 31, 2020 |  | 91,310,000 |  |  |
Conversion benefit included in Convertible Notes issued | 114 |  | 114 |  |
Issuance of common stock under equity incentive plan | 883 | $ 1 | 882 |  |
Issuance of common stock under equity incentive plan (in shares) |  | 97,000 |  |  |
Stock-based compensation expense | 11,486 |  | 11,486 |  |
Net loss | (52,614) |  |  |  | (52,614)
Net unrealized gain (loss) on short-term investments | (136) |  |  | (136) |
Comprehensive loss | (52,750) |  |  |  |
Balance at Mar. 31, 2021 | 196,225 | $ 914 | 1,640,552 | 121 | (1,445,362)
Balance (in shares) at Mar. 31, 2021 |  | 91,407,000 |  |  |
Balance at Dec. 31, 2020 | $ 236,492 | $ 913 | 1,628,070 | 257 | (1,392,748)
Balance (in shares) at Dec. 31, 2020 |  | 91,310,000 |  |  |
Issuance of common stock on exercise of stock options (in shares) | 357,839 |  |  |  |
Net loss | $ (113,629) |  |  |  |
Net unrealized gain (loss) on short-term investments | (216) |  |  |  |
Comprehensive loss | (113,845) |  |  |  |
Balance at Jun. 30, 2021 | 160,112 | $ 1,019 | 1,665,429 | 41 | (1,506,377)
Balance (in shares) at Jun. 30, 2021 |  | 101,874,000 |  |  |
Balance at Mar. 31, 2021 | 196,225 | $ 914 | 1,640,552 | 121 | (1,445,362)
Balance (in shares) at Mar. 31, 2021 |  | 91,407,000 |  |  |
Conversion benefit included in Convertible Notes issued | 116 |  | 116 |  |
Issuance of common stock under equity incentive plan | 4,218 | $ 4 | 4,214 |  |
Issuance of common stock under equity incentive plan (in shares) |  | 313,000 |  |  |
Issuance of common stock on under the employee stock purchase plan | 1,529 | $ 1 | 1,528 |  |
Issuance of common stock on under the employee stock purchase plan (in shares) |  | 110,000 |  |  |
Issuance of common stock on exercise of warrants |  | $ 2 | (2) |  |
Issuance of common stock on exercise of warrants (in shares) |  | 195,000 |  |  |
Issuance of common stock on conversion of notes | 7,878 | $ 98 | 7,780 |  |
Issuance of common stock on conversion of notes (in shares) |  | 9,849,000 |  |  |
Stock-based compensation expense | 11,241 |  | 11,241 |  |
Net loss | (61,015) |  |  |  | (61,015)
Net unrealized gain (loss) on short-term investments | (80) |  |  | (80) |
Comprehensive loss | (61,095) |  |  |  |
Balance at Jun. 30, 2021 | $ 160,112 | $ 1,019 | $ 1,665,429 | $ 41 | $ (1,506,377)
Balance (in shares) at Jun. 30, 2021 |  | 101,874,000 |  |  |

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