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Condensed Consolidated Statements of Stockholders' Equity (Unaudited) - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-In Capital
Accumulated Other Comprehensive Income
Accumulated Deficit
Balance at Dec. 31, 2019 $ 403,835 $ 903 $ 1,568,317 $ 85 $ (1,165,470)
Balance (in shares) at Dec. 31, 2019   90,304,000      
Conversion benefit included in Convertible Notes issued 108   108    
Issuance of common stock under equity incentive plan 504   504    
Issuance of common stock under equity incentive plan (in shares)   70,000      
Issuance of common stock on exercise of warrants 3 $ 3      
Issuance of common stock on exercise of warrants (in shares)   268,000      
Stock-based compensation expense 11,974   11,974    
Net loss (51,579)       (51,579)
Net unrealized gain (loss) on short-term investments 623     623  
Comprehensive loss (50,956)        
Balance at Mar. 31, 2020 365,468 $ 906 1,580,903 708 (1,217,049)
Balance (in shares) at Mar. 31, 2020   90,642,000      
Balance at Dec. 31, 2019 403,835 $ 903 1,568,317 85 (1,165,470)
Balance (in shares) at Dec. 31, 2019   90,304,000      
Net loss (106,769)        
Net unrealized gain (loss) on short-term investments 868        
Comprehensive loss (105,901)        
Balance at Jun. 30, 2020 324,058 $ 908 1,594,436 953 (1,272,239)
Balance (in shares) at Jun. 30, 2020   90,820,000      
Balance at Mar. 31, 2020 365,468 $ 906 1,580,903 708 (1,217,049)
Balance (in shares) at Mar. 31, 2020   90,642,000      
Conversion benefit included in Convertible Notes issued 109   109    
Issuance of common stock under equity incentive plan 805 $ 1 804    
Issuance of common stock under equity incentive plan (in shares)   54,000      
Issuance of common stock on under the employee stock purchase plan 1,507 $ 1 1,506    
Issuance of common stock on under the employee stock purchase plan (in shares)   124,000      
Stock-based compensation expense 11,114   11,114    
Net loss (55,190)       (55,190)
Net unrealized gain (loss) on short-term investments 245     245  
Comprehensive loss (54,945)        
Balance at Jun. 30, 2020 324,058 $ 908 1,594,436 953 (1,272,239)
Balance (in shares) at Jun. 30, 2020   90,820,000      
Balance at Dec. 31, 2020 236,492 $ 913 1,628,070 257 (1,392,748)
Balance (in shares) at Dec. 31, 2020   91,310,000      
Conversion benefit included in Convertible Notes issued 114   114    
Issuance of common stock under equity incentive plan 883 $ 1 882    
Issuance of common stock under equity incentive plan (in shares)   97,000      
Stock-based compensation expense 11,486   11,486    
Net loss (52,614)       (52,614)
Net unrealized gain (loss) on short-term investments (136)     (136)  
Comprehensive loss (52,750)        
Balance at Mar. 31, 2021 196,225 $ 914 1,640,552 121 (1,445,362)
Balance (in shares) at Mar. 31, 2021   91,407,000      
Balance at Dec. 31, 2020 $ 236,492 $ 913 1,628,070 257 (1,392,748)
Balance (in shares) at Dec. 31, 2020   91,310,000      
Issuance of common stock on exercise of stock options (in shares) 357,839        
Net loss $ (113,629)        
Net unrealized gain (loss) on short-term investments (216)        
Comprehensive loss (113,845)        
Balance at Jun. 30, 2021 160,112 $ 1,019 1,665,429 41 (1,506,377)
Balance (in shares) at Jun. 30, 2021   101,874,000      
Balance at Mar. 31, 2021 196,225 $ 914 1,640,552 121 (1,445,362)
Balance (in shares) at Mar. 31, 2021   91,407,000      
Conversion benefit included in Convertible Notes issued 116   116    
Issuance of common stock under equity incentive plan 4,218 $ 4 4,214    
Issuance of common stock under equity incentive plan (in shares)   313,000      
Issuance of common stock on under the employee stock purchase plan 1,529 $ 1 1,528    
Issuance of common stock on under the employee stock purchase plan (in shares)   110,000      
Issuance of common stock on exercise of warrants   $ 2 (2)    
Issuance of common stock on exercise of warrants (in shares)   195,000      
Issuance of common stock on conversion of notes 7,878 $ 98 7,780    
Issuance of common stock on conversion of notes (in shares)   9,849,000      
Stock-based compensation expense 11,241   11,241    
Net loss (61,015)       (61,015)
Net unrealized gain (loss) on short-term investments (80)     (80)  
Comprehensive loss (61,095)        
Balance at Jun. 30, 2021 $ 160,112 $ 1,019 $ 1,665,429 $ 41 $ (1,506,377)
Balance (in shares) at Jun. 30, 2021   101,874,000