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Consolidated Statements of Stockholders' Equity (Deficit) - USD ($) shares in Thousands, $ in Thousands | Total | Private Placement | Common Stock | Common Stock Private Placement | Additional Paid-In Capital | Additional Paid-In Capital Private Placement | Accumulated Other Comprehensive Income (Loss) | Accumulated Deficit
--- | --- | --- | --- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2021 | $ 77,570 |  | $ 1,020 |  | $ 1,689,987 |  | $ (6) | $ (1,613,431)
Balance (in shares) at Dec. 31, 2021 |  |  | 102,005 |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 1,444 |  | $ 4 |  | 1,440 |  |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  |  | 407 |  |  |  |  |
Issuance of common stock under equity incentive plan | (1,530) |  | $ 6 |  | (1,536) |  |  |
Issuance of common stock under equity incentive plan (in shares) |  |  | 614 |  |  |  |  |
Issuance of common stock in a private placement |  | $ 75,145 |  | $ 161 |  | $ 74,984 |  |
Issuance of common stock in a private placement (in shares) |  |  |  | 16,129 |  |  |  |
Stock-based compensation expense | 42,980 |  |  |  | 42,980 |  |  |
Net loss | (182,024) |  |  |  |  |  |  | (182,024)
Net unrealized gain (loss) on short-term investments | (13) |  |  |  |  |  | (13) |
Comprehensive loss | (182,037) |  |  |  |  |  |  |
Balance at Dec. 31, 2022 | 13,572 |  | $ 1,191 |  | 1,807,855 |  | (19) | (1,795,455)
Balance (in shares) at Dec. 31, 2022 |  |  | 119,155 |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 904 |  | $ 7 |  | 897 |  |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  |  | 717 |  |  |  |  |
Issuance of common stock under equity incentive plan | (903) |  | $ 11 |  | (914) |  |  |
Issuance of common stock under equity incentive plan (in shares) |  |  | 1,130 |  |  |  |  |
Issuance of common stock in a private placement |  | $ 29,755 |  | $ 208 |  | $ 29,547 |  |
Issuance of common stock in a private placement (in shares) |  |  |  | 20,735 |  |  |  |
Issuance of common stock on exercise of pre-funded warrants | 1 |  | $ 86 |  | (85) |  |  |
Issuance of common stock on exercise of pre-funded warrants (in shares) |  |  | 8,548 |  |  |  |  |
Issuance of warrants in debt financing | 371 |  |  |  | 371 |  |  |
Stock-based compensation expense | 32,854 |  |  |  | 32,854 |  |  |
Net loss | (110,559) |  |  |  |  |  |  | (110,559)
Net unrealized gain (loss) on short-term investments | 32 |  |  |  |  |  | 32 |
Comprehensive loss | (110,527) |  |  |  |  |  |  |
Balance at Dec. 31, 2023 | (33,973) |  | $ 1,503 |  | 1,870,525 |  | 13 | (1,906,014)
Balance (in shares) at Dec. 31, 2023 |  |  | 150,285 |  |  |  |  |
Issuance of common stock under Employee Stock Purchase Plan | 637 |  | $ 6 |  | 631 |  |  |
Issuance of common stock under Employee Stock Purchase Plan (in shares) |  |  | 640 |  |  |  |  |
Issuance of common stock under equity incentive plan | 303 |  | $ 12 |  | 291 |  |  |
Issuance of common stock under equity incentive plan (in shares) |  |  | 1,203 |  |  |  |  |
Stock-based compensation expense | 12,962 |  |  |  | 12,962 |  |  |
Net loss | (13,580) |  |  |  |  |  |  | (13,580)
Comprehensive loss | (13,580) |  |  |  |  |  |  |
Balance at Dec. 31, 2024 | $ (33,651) |  | $ 1,521 |  | $ 1,884,409 |  | $ 13 | $ (1,919,594)
Balance (in shares) at Dec. 31, 2024 |  |  | 152,128 |  |  |  |  |

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