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Collaborative Arrangements - Strategic Collaboration and License Agreement (Details) - USD ($) $ in Thousands | 12 Months Ended |
--- | --- | ---
Dec. 31, 2023 | Dec. 31, 2022 | Dec. 31, 2021 | Mar. 31, 2020
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |
Contract liabilities: deferred revenue | $ 10,761 | $ 30,609 |  |
Milestone Payment Per Target Gene |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |
Contingent receivable |  |  |  | $ 104,000
Takeda |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |
Contract liabilities: deferred revenue | 0 | 900 |  |
Revenue recognized, including opening balance | $ 2,000 | $ 4,900 | $ 7,400 |
Takeda | Up-front Payment |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |
Contract liabilities: deferred revenue |  |  |  | $ 8,500

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