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Collaborative Arrangements - Master Collaboration and Research Agreement, Stock Purchase Agreement and Enzyme Supply Agreement (Details) - USD ($) $ in Thousands | 1 Months Ended | 12 Months Ended
--- | --- | ---
Jul. 31, 2022 | Dec. 31, 2021 | Jun. 30, 2020 | Dec. 31, 2023 | Dec. 31, 2022 | Dec. 31, 2021
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Investments in non-marketable securities |  |  |  | $ 1,191 | $ 5,300 | $ 7,630
Contract liabilities: deferred revenue |  |  |  | 10,761 | 30,609 |
Related Party |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Investments in non-marketable securities |  |  |  | 0 | 0 | 7,630
MAI | Commercialization And Enzyme Supply Agreement |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Aggregate commercial sales, milestone |  |  | $ 5,000 |  |  |
MAI | Master Collaboration & Research Agreement |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Revenue recognized from transactions |  |  |  |  | 1,200 | $ 2,000
MAI | Revenue sharing arrangement | Commercialization And Enzyme Supply Agreement |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Contract liabilities: deferred revenue |  |  | $ 500 |  |  |
MAI | Product revenue | Master Collaboration & Research Agreement |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Revenue recognized from transactions |  |  |  | $ 200 | $ 500 |
MAI | Series A Preferred Stock |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Investment owned, balance (in shares) |  |  | 1,587,050 |  |  |
Investments in non-marketable securities |  |  | $ 1,000 |  |  |
MAI | Series B Preferred Stock | Related Party |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Milestone payments received (in shares) | 1,587,049 | 1,587,049 |  |  |  |
MAI | Series A and B Preferred Stock | Master Collaboration & Research Agreement |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
Investment owned, balance (in shares) |  | 3,491,505 |  |  | 1,587,049 | 3,491,505
MAI Agreement | Related Party |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |
MAI agreement milestone payment received | $ 1,000 | $ 1,000 |  |  |  |

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