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Collaborative Arrangements - Enzyme Supply Agreement (Details) - USD ($) $ in Thousands | 12 Months Ended
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Dec. 31, 2023 | Dec. 31, 2022 | Dec. 31, 2021
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |
Revenues | $ 70,143 | $ 138,590 | $ 104,754
Contract liabilities: deferred revenue | 10,761 | 30,609 |
Supply Agreement |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |
Contract liabilities: deferred revenue | 0 | 3,300 |
Product revenue |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |
Revenues | 42,906 | $ 116,676 | $ 70,657
Product revenue | Enzyme Supply Agreement |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |
Revenues | 3,200 |  |
Revenue from contract with customer, settlement fee | $ 1,300 |  |

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