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Collaborative Arrangements - Enzyme Supply Agreement (Details) - USD ($)
$ in Thousands
 12 Months Ended
Dec. 31, 2023
Dec. 31, 2022
Dec. 31, 2021
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]      
Revenues $ 70,143 $ 138,590 $ 104,754
Contract liabilities: deferred revenue 10,761 30,609  
Supply Agreement      
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]      
Contract liabilities: deferred revenue 0 3,300  
Product revenue      
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]      
Revenues 42,906 $ 116,676 $ 70,657
Product revenue | Enzyme Supply Agreement      
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]      
Revenues 3,200    
Revenue from contract with customer, settlement fee $ 1,300