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Collaborative Arrangements - Master Collaboration and Research Agreement, Stock Purchase Agreement and Enzyme Supply Agreement (Details) - USD ($) $ in Thousands	1 Months Ended			12 Months Ended
	Jul. 31, 2022	Dec. 31, 2021	Jun. 30, 2020	Dec. 31, 2023	Dec. 31, 2022	Dec. 31, 2021
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Investments in non-marketable securities				$ 1,191	$ 5,300	$ 7,630
Contract liabilities: deferred revenue				10,761	30,609
Related Party
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Investments in non-marketable securities				0	0	7,630
MAI | Commercialization And Enzyme Supply Agreement
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Aggregate commercial sales, milestone			$ 5,000
MAI | Master Collaboration & Research Agreement
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenue recognized from transactions					1,200	$ 2,000
MAI | Revenue sharing arrangement | Commercialization And Enzyme Supply Agreement
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Contract liabilities: deferred revenue			$ 500
MAI | Product revenue | Master Collaboration & Research Agreement
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Revenue recognized from transactions				$ 200	$ 500
MAI | Series A Preferred Stock
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Investment owned, balance (in shares)			1,587,050
Investments in non-marketable securities			$ 1,000
MAI | Series B Preferred Stock | Related Party
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Milestone payments received (in shares)	1,587,049	1,587,049
MAI | Series A and B Preferred Stock | Master Collaboration & Research Agreement
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Investment owned, balance (in shares)		3,491,505			1,587,049	3,491,505
MAI Agreement | Related Party
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
MAI agreement milestone payment received	$ 1,000	$ 1,000

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