<SEC-DOCUMENT>0001716774-25-000007.txt : 20251107
<SEC-HEADER>0001716774-25-000007.hdr.sgml : 20251107
<ACCEPTANCE-DATETIME>20251107112810
ACCESSION NUMBER:		0001716774-25-000007
CONFORMED SUBMISSION TYPE:	SCHEDULE 13G/A
PUBLIC DOCUMENT COUNT:		1
FILED AS OF DATE:		20251107
DATE AS OF CHANGE:		20251107

SUBJECT COMPANY:	

	COMPANY DATA:	
		COMPANY CONFORMED NAME:			CODEXIS, INC.
		CENTRAL INDEX KEY:			0001200375
		STANDARD INDUSTRIAL CLASSIFICATION:	INDUSTRIAL ORGANIC CHEMICALS [2860]
		ORGANIZATION NAME:           	08 Industrial Applications and Services
		EIN:				710872999
		STATE OF INCORPORATION:			DE
		FISCAL YEAR END:			1231

	FILING VALUES:
		FORM TYPE:		SCHEDULE 13G/A
		SEC ACT:		1934 Act
		SEC FILE NUMBER:	005-85965
		FILM NUMBER:		251461249

	BUSINESS ADDRESS:	
		STREET 1:		200 PENOBSCOT DRIVE
		CITY:			REDWOOD CITY
		STATE:			CA
		ZIP:			94063
		BUSINESS PHONE:		650-421-8100

	MAIL ADDRESS:	
		STREET 1:		200 PENOBSCOT DRIVE
		CITY:			REDWOOD CITY
		STATE:			CA
		ZIP:			94063

	FORMER COMPANY:	
		FORMER CONFORMED NAME:	CODEXIS INC
		DATE OF NAME CHANGE:	20021022

FILED BY:		

	COMPANY DATA:	
		COMPANY CONFORMED NAME:			Aberdeen Group plc
		CENTRAL INDEX KEY:			0001716774
		ORGANIZATION NAME:           	
		EIN:				000000000
		STATE OF INCORPORATION:			X0
		FISCAL YEAR END:			1231

	FILING VALUES:
		FORM TYPE:		SCHEDULE 13G/A

	BUSINESS ADDRESS:	
		STREET 1:		1 GEORGE STREET
		CITY:			EDINBURGH
		STATE:			X0
		ZIP:			EH2 2LL
		BUSINESS PHONE:		0131 245 2552

	MAIL ADDRESS:	
		STREET 1:		1 GEORGE STREET
		CITY:			EDINBURGH
		STATE:			X0
		ZIP:			EH2 2LL

	FORMER COMPANY:	
		FORMER CONFORMED NAME:	abrdn plc
		DATE OF NAME CHANGE:	20210707

	FORMER COMPANY:	
		FORMER CONFORMED NAME:	Standard Life Aberdeen plc
		DATE OF NAME CHANGE:	20170912
</SEC-HEADER>
<DOCUMENT>
<TYPE>SCHEDULE 13G/A
<SEQUENCE>1
<FILENAME>primary_doc.xml
<TEXT>
<XML>
<?xml version="1.0" encoding="UTF-8"?><edgarSubmission xmlns="http://www.sec.gov/edgar/schedule13g" xmlns:ns1="http://www.sec.gov/edgar/common">
  <headerData>
    <submissionType>SCHEDULE 13G/A</submissionType>
    <previousAccessionNumber>0001085146-25-004305</previousAccessionNumber>
    <filerInfo>
      <filer>
        <filerCredentials>
          <cik>0001716774</cik>
          <ccc>XXXXXXXX</ccc>
        </filerCredentials>
      </filer>
      <liveTestFlag>LIVE</liveTestFlag>



    </filerInfo>
  </headerData>
  <formData>
    <coverPageHeader>
      <amendmentNo>1</amendmentNo>
      <securitiesClassTitle>Common Stock, par value $0.0001 per share</securitiesClassTitle>
      <eventDateRequiresFilingThisStatement>10/01/2025</eventDateRequiresFilingThisStatement>
      <issuerInfo>
        <issuerCik>0001200375</issuerCik>
        <issuerName>CODEXIS, INC.</issuerName>
        <issuerCusip>192005106</issuerCusip>
        <issuerPrincipalExecutiveOfficeAddress>
          <ns1:street1>200 PENOBSCOT DRIVE</ns1:street1>
          <ns1:city>REDWOOD CITY</ns1:city>
          <ns1:stateOrCountry>CA</ns1:stateOrCountry>
          <ns1:zipCode>94063</ns1:zipCode>
        </issuerPrincipalExecutiveOfficeAddress>
      </issuerInfo>
      <designateRulesPursuantThisScheduleFiled>
        <designateRulePursuantThisScheduleFiled>Rule 13d-1(b)</designateRulePursuantThisScheduleFiled>
      </designateRulesPursuantThisScheduleFiled>
    </coverPageHeader>
    <coverPageHeaderReportingPersonDetails>


      <reportingPersonName>Aberdeen Group plc</reportingPersonName>
      <citizenshipOrOrganization>X0</citizenshipOrOrganization>
      <reportingPersonBeneficiallyOwnedNumberOfShares>
        <soleVotingPower>0</soleVotingPower>
        <sharedVotingPower>4092062</sharedVotingPower>
        <soleDispositivePower>0</soleDispositivePower>
        <sharedDispositivePower>4092062</sharedDispositivePower>
      </reportingPersonBeneficiallyOwnedNumberOfShares>
      <reportingPersonBeneficiallyOwnedAggregateNumberOfShares>4092062</reportingPersonBeneficiallyOwnedAggregateNumberOfShares>
      <aggregateAmountExcludesCertainSharesFlag>N</aggregateAmountExcludesCertainSharesFlag>
      <classPercent>4.53</classPercent>
      <typeOfReportingPerson>HC</typeOfReportingPerson>
    </coverPageHeaderReportingPersonDetails>
    <coverPageHeaderReportingPersonDetails>


      <reportingPersonName>abrdn Inc.</reportingPersonName>
      <citizenshipOrOrganization>X1</citizenshipOrOrganization>
      <reportingPersonBeneficiallyOwnedNumberOfShares>
        <soleVotingPower>0</soleVotingPower>
        <sharedVotingPower>4092062</sharedVotingPower>
        <soleDispositivePower>0</soleDispositivePower>
        <sharedDispositivePower>4092062</sharedDispositivePower>
      </reportingPersonBeneficiallyOwnedNumberOfShares>
      <reportingPersonBeneficiallyOwnedAggregateNumberOfShares>4092062</reportingPersonBeneficiallyOwnedAggregateNumberOfShares>
      <aggregateAmountExcludesCertainSharesFlag>N</aggregateAmountExcludesCertainSharesFlag>
      <classPercent>4.53</classPercent>
      <typeOfReportingPerson>CO</typeOfReportingPerson>
      <typeOfReportingPerson>IA</typeOfReportingPerson>
    </coverPageHeaderReportingPersonDetails>
    <items>
      <item1>
        <issuerName>CODEXIS, INC.</issuerName>
        <issuerPrincipalExecutiveOfficeAddress>200 PENOBSCOT DRIVE, REDWOOD CITY, CALIFORNIA
94063</issuerPrincipalExecutiveOfficeAddress>
      </item1>
      <item2>
        <filingPersonName>Aberdeen Group plc

abrdn Inc.</filingPersonName>
        <principalBusinessOfficeOrResidenceAddress>Aberdeen Group plc:
1 George Street
Edinburgh, United Kingdom
EH2 2LL

abrdn Inc.
1900 Market Street Suite 200.
Philadelphia,
PA 19103</principalBusinessOfficeOrResidenceAddress>
        <citizenship>Aberdeen Group plc - UNITED KINGDOM

abrdn Inc. - UNITED STATES</citizenship>
      </item2>
      <item3>
        <notApplicableFlag>N</notApplicableFlag>
        <typeOfPersonFiling>HC</typeOfPersonFiling>
      </item3>
      <item4>
        <amountBeneficiallyOwned>4,092,062</amountBeneficiallyOwned>
        <classPercent>4.53</classPercent>
        <numberOfSharesPersonHas>
          <solePowerOrDirectToVote>Aberdeen Group plc - 0

abrdn Inc. - 0</solePowerOrDirectToVote>
          <sharedPowerOrDirectToVote>Aberdeen Group plc - 4,092,062

abrdn Inc. - 4,092,062</sharedPowerOrDirectToVote>
          <solePowerOrDirectToDispose>Aberdeen Group plc - 0

abrdn Inc. - 0</solePowerOrDirectToDispose>
          <sharedPowerOrDirectToDispose>Aberdeen Group plc - 4,092,062

abrdn Inc. - 4,092,062</sharedPowerOrDirectToDispose>
        </numberOfSharesPersonHas>
      </item4>
      <item5>
        <notApplicableFlag>N</notApplicableFlag>
        <classOwnership5PercentOrLess>Y</classOwnership5PercentOrLess>
      </item5>
      <item6>
        <notApplicableFlag>Y</notApplicableFlag>
      </item6>
      <item7>
        <notApplicableFlag>N</notApplicableFlag>
        <subsidiaryIdentificationAndClassification>See Exhibit</subsidiaryIdentificationAndClassification>
      </item7>
      <item8>
        <notApplicableFlag>Y</notApplicableFlag>
      </item8>
      <item9>
        <notApplicableFlag>Y</notApplicableFlag>
      </item9>
      <item10>
        <notApplicableFlag>N</notApplicableFlag>
        <certifications>By signing below I certify that, to the best of my knowledge and belief, the securities referred to above were acquired and are held in the ordinary course of business and were not acquired and are not held for the purpose of or with the effect of changing or influencing the control of the issuer of the securities and were not acquired and are not held in connection with or as a participant in any transaction having that purpose or effect, other than activities solely in connection with a nomination under ?? 240.14a-11.</certifications>
      </item10>
    </items>
    <exhibitInfo>Aberdeen Group plc
abrdn Holdings Limited
abrdn Inc.

Aberdeen Group plc is the parent company.
abrdn Holdings Limited is the intermediate holding company for abrdn Inc.
abrdn Inc. beneficially owns on behalf of our underlying clients less than 5% of the outstanding shares of the security class being reported on this Schedule 13G.</exhibitInfo>
    <signatureInformation>
      <reportingPersonName>Aberdeen Group plc</reportingPersonName>
      <signatureDetails>
        <signature>Jenny Clarke</signature>
        <title>Major Shareholding Reporting Analyst</title>
        <date>10/29/2025</date>
      </signatureDetails>
    </signatureInformation>
    <signatureInformation>
      <reportingPersonName>abrdn Inc.</reportingPersonName>
      <signatureDetails>
        <signature>Jenny Clarke</signature>
        <title>Major Shareholding Reporting Analyst</title>
        <date>10/29/2025</date>
      </signatureDetails>
    </signatureInformation>
  </formData>
</edgarSubmission>
</XML>
</TEXT>
</DOCUMENT>
</SEC-DOCUMENT>
