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Consolidated Statements of Stockholders' Equity (USD $) In Thousands, except Share data, unless otherwise specified | Total | Common Stock [Member] | Additional Paid-in Capital [Member] | Accumulated Deficit [Member] | Accumulated Other Comprehensive Income/ (Loss) [Member]
--- | --- | --- | --- | --- | ---
Beginning Balances at Dec. 31, 2011 | $ 80,132 us-gaap_StockholdersEquity | $ 526 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | $ 332,839 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember | $ (253,245)us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_RetainedEarningsMember | $ 12 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AccumulatedOtherComprehensiveIncomeMember
Beginning Balances, shares at Dec. 31, 2011 |  | 52,554,795 us-gaap_SharesOutstanding / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units | 1,219 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation | 3 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 1,216 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, shares |  | 328,355 us-gaap_StockIssuedDuringPeriodSharesShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock under employee stock purchase plan | 457 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan | 2 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 455 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock under employee stock purchase plan, shares | 175,375 us-gaap_StockIssuedDuringPeriodSharesEmployeeStockPurchasePlans | 175,375 us-gaap_StockIssuedDuringPeriodSharesEmployeeStockPurchasePlans / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Stock-based compensation | 5,338 us-gaap_AdjustmentsToAdditionalPaidInCapitalSharebasedCompensationRequisiteServicePeriodRecognitionValue |  | 5,338 us-gaap_AdjustmentsToAdditionalPaidInCapitalSharebasedCompensationRequisiteServicePeriodRecognitionValue / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Comprehensive loss: |  |  |  |  |
Net unrealized gain (loss) on marketable securities | 14 us-gaap_OtherComprehensiveIncomeUnrealizedHoldingGainLossOnSecuritiesArisingDuringPeriodNetOfTax |  |  |  | 14 us-gaap_OtherComprehensiveIncomeUnrealizedHoldingGainLossOnSecuritiesArisingDuringPeriodNetOfTax / us-gaap_StatementEquityComponentsAxis = us-gaap_AccumulatedOtherComprehensiveIncomeMember
Net loss | (22,264)us-gaap_NetIncomeLoss |  |  | (22,264)us-gaap_NetIncomeLoss / us-gaap_StatementEquityComponentsAxis = us-gaap_RetainedEarningsMember |
Comprehensive loss | (22,250)us-gaap_ComprehensiveIncomeNetOfTax |  |  |  |
Ending Balances at Dec. 31, 2012 | 64,896 us-gaap_StockholdersEquity | 531 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 339,848 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember | (275,509)us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_RetainedEarningsMember | 26 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AccumulatedOtherComprehensiveIncomeMember
Ending Balances, shares at Dec. 31, 2012 |  | 53,058,525 us-gaap_SharesOutstanding / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock in connection with underwritten public offering, net of issuance costs | 69,492 us-gaap_StockIssuedDuringPeriodValueNewIssues | 70 us-gaap_StockIssuedDuringPeriodValueNewIssues / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 69,422 us-gaap_StockIssuedDuringPeriodValueNewIssues / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock in connection with underwritten public offering, net of issuance costs, shares |  | 7,015,000 us-gaap_StockIssuedDuringPeriodSharesNewIssues / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units | 6,118 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation | 19 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 6,099 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, shares |  | 1,889,818 us-gaap_StockIssuedDuringPeriodSharesShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock under employee stock purchase plan | 496 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan | 1 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 495 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock under employee stock purchase plan, shares | 180,551 us-gaap_StockIssuedDuringPeriodSharesEmployeeStockPurchasePlans | 180,551 us-gaap_StockIssuedDuringPeriodSharesEmployeeStockPurchasePlans / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock in connection with acquisition of Ceregene, Inc. | 1,200 us-gaap_StockIssuedDuringPeriodValueAcquisitions | 1 us-gaap_StockIssuedDuringPeriodValueAcquisitions / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 1,199 us-gaap_StockIssuedDuringPeriodValueAcquisitions / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock in connection with acquisition of Ceregene, Inc., shares |  | 99,998 us-gaap_StockIssuedDuringPeriodSharesAcquisitions / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Stock-based compensation | 6,146 us-gaap_AdjustmentsToAdditionalPaidInCapitalSharebasedCompensationRequisiteServicePeriodRecognitionValue |  | 6,146 us-gaap_AdjustmentsToAdditionalPaidInCapitalSharebasedCompensationRequisiteServicePeriodRecognitionValue / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Comprehensive loss: |  |  |  |  |
Net unrealized gain (loss) on marketable securities | (14)us-gaap_OtherComprehensiveIncomeUnrealizedHoldingGainLossOnSecuritiesArisingDuringPeriodNetOfTax |  |  |  | (14)us-gaap_OtherComprehensiveIncomeUnrealizedHoldingGainLossOnSecuritiesArisingDuringPeriodNetOfTax / us-gaap_StatementEquityComponentsAxis = us-gaap_AccumulatedOtherComprehensiveIncomeMember
Net loss | (26,624)us-gaap_NetIncomeLoss |  |  | (26,624)us-gaap_NetIncomeLoss / us-gaap_StatementEquityComponentsAxis = us-gaap_RetainedEarningsMember |
Comprehensive loss | (26,638)us-gaap_ComprehensiveIncomeNetOfTax |  |  |  |
Ending Balances at Dec. 31, 2013 | 121,710 us-gaap_StockholdersEquity | 622 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 423,209 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember | (302,133)us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_RetainedEarningsMember | 12 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AccumulatedOtherComprehensiveIncomeMember
Ending Balances, shares at Dec. 31, 2013 |  | 62,243,892 us-gaap_SharesOutstanding / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock in connection with underwritten public offering, net of issuance costs | 93,796 us-gaap_StockIssuedDuringPeriodValueNewIssues | 44 us-gaap_StockIssuedDuringPeriodValueNewIssues / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 93,752 us-gaap_StockIssuedDuringPeriodValueNewIssues / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock in connection with underwritten public offering, net of issuance costs, shares |  | 4,444,444 us-gaap_StockIssuedDuringPeriodSharesNewIssues / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units | 7,533 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation | 23 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 7,510 us-gaap_StockIssuedDuringPeriodValueShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, shares |  | 2,266,855 us-gaap_StockIssuedDuringPeriodSharesShareBasedCompensation / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Issuance of common stock under employee stock purchase plan | 848 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan | 1 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | 847 us-gaap_StockIssuedDuringPeriodValueEmployeeStockPurchasePlan / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Issuance of common stock under employee stock purchase plan, shares | 107,203 us-gaap_StockIssuedDuringPeriodSharesEmployeeStockPurchasePlans | 107,203 us-gaap_StockIssuedDuringPeriodSharesEmployeeStockPurchasePlans / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |
Stock-based compensation | 9,200 us-gaap_AdjustmentsToAdditionalPaidInCapitalSharebasedCompensationRequisiteServicePeriodRecognitionValue |  | 9,200 us-gaap_AdjustmentsToAdditionalPaidInCapitalSharebasedCompensationRequisiteServicePeriodRecognitionValue / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember |  |
Comprehensive loss: |  |  |  |  |
Net unrealized gain (loss) on marketable securities | (37)us-gaap_OtherComprehensiveIncomeUnrealizedHoldingGainLossOnSecuritiesArisingDuringPeriodNetOfTax |  |  |  | (37)us-gaap_OtherComprehensiveIncomeUnrealizedHoldingGainLossOnSecuritiesArisingDuringPeriodNetOfTax / us-gaap_StatementEquityComponentsAxis = us-gaap_AccumulatedOtherComprehensiveIncomeMember
Net loss | (26,417)us-gaap_NetIncomeLoss |  |  | (26,417)us-gaap_NetIncomeLoss / us-gaap_StatementEquityComponentsAxis = us-gaap_RetainedEarningsMember |
Comprehensive loss | (26,454)us-gaap_ComprehensiveIncomeNetOfTax |  |  |  |
Ending Balances at Dec. 31, 2014 | $ 206,633 us-gaap_StockholdersEquity | $ 690 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember | $ 534,518 us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AdditionalPaidInCapitalMember | $ (328,550)us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_RetainedEarningsMember | $ (25)us-gaap_StockholdersEquity / us-gaap_StatementEquityComponentsAxis = us-gaap_AccumulatedOtherComprehensiveIncomeMember
Ending Balances, shares at Dec. 31, 2014 |  | 69,062,394 us-gaap_SharesOutstanding / us-gaap_StatementEquityComponentsAxis = us-gaap_CommonStockMember |  |  |

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