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Major Customers, Partnerships and Strategic Alliances - Additional Information (Detail) | 1 Months Ended | 12 Months Ended |
--- | --- | --- | ---
Jan. 31, 2014 USD ($) Program | Jun. 30, 2012 Targets | Jan. 31, 2012 USD ($) Targets | Dec. 31, 2015 USD ($) Product Targets | Dec. 31, 2014 USD ($) | Dec. 31, 2013 USD ($) | Sep. 01, 2015 USD ($)
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |
Number of research programs | Program | 2 |  |  |  |  |  |
Research and development |  |  |  | $ 67,198,000 | $ 56,974,000 | $ 37,039,000 |
Biogen Inc [Member] |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |
Revenues under agreement | $ 20,000,000 |  |  |  |  |  |
Potential amount eligible to receive for certain milestones | 126,300,000 |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones | 167,500,000 |  |  |  |  |  |
Milestone revenue receivable | 293,800,000 |  |  |  |  |  |
Milestone payments received |  |  |  | $ 0 |  |  |
Number of products approved | Product |  |  |  | 0 |  |  |
Royalty revenues |  |  |  | $ 0 |  |  |
Research program to develop laboratory research reagents |  |  |  | 40 months |  |  |
Deferred revenue |  |  |  | $ 8,600,000 |  |  |
Biogen Inc [Member] | Sickle cell disease [Member] |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |
Milestone revenue receivable | 7,500,000 |  |  |  |  |  |
Biogen Inc [Member] | Beta-thalassemia Project [Member] |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |
Milestone revenue receivable | $ 7,500,000 |  |  |  |  |  |
Research and development |  |  |  | 6,500,000 | 5,200,000 |  |
Biogen Inc [Member] | Other Projects [Member] |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |
Research and development |  |  |  | $ 2,900,000 | 3,500,000 |  |
Shire AG [Member] |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |
Revenues under agreement |  |  | $ 13,000,000 |  |  |  |
Research program to develop laboratory research reagents |  |  |  | 6 years |  |  |
Deferred revenue |  |  |  | $ 5,100,000 |  |  |
Research and development |  |  |  | $ 13,500,000 | 21,100,000 | $ 14,200,000 |
Aggregate number of gene targets | Targets |  |  | 7 |  |  |  |
Number of initial gene targets | Targets |  |  | 4 |  |  |  |
Number of gene targets | Targets |  | 5 |  |  |  |  |
Number of additional gene targets | Targets |  | 2 |  | 1 |  |  |
Recognition of milestone |  |  |  |  | $ 1,000,000 |  |
Amount of obligation related to reimbursement |  |  |  |  |  |  | $ 4,000,000
Revenue recognized related to prior obligations |  |  |  | $ 3,400,000 |  |  |

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