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Major Customers, Partnerships and Strategic Alliances - Additional Information (Detail)		1 Months Ended				3 Months Ended		9 Months Ended		12 Months Ended
	Sep. 01, 2015 USD ($)	Jan. 31, 2016 USD ($)	Jan. 31, 2014 USD ($) Program	Jun. 30, 2012 Targets	Jan. 31, 2012 USD ($) Targets	Sep. 30, 2016 USD ($)	Sep. 30, 2015 USD ($)	Sep. 30, 2016 USD ($) Product Targets	Sep. 30, 2015 USD ($)	Dec. 31, 2014 USD ($)
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Number of research programs | Program			2
Research and development						$ 17,008,000	$ 16,694,000	$ 51,728,000	$ 47,292,000
Biogen Inc [Member]
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Revenues under agreement			$ 20,000,000
Milestone payments received								$ 0
Number of products approved | Product								0
Royalty revenues								$ 0
Deferred revenue						6,700,000		6,700,000
Biogen Inc [Member] | Beta-thalassemia Project [Member]
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Research and development						1,200,000	2,300,000	5,300,000	4,700,000
Biogen Inc [Member] | Other Projects [Member]
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Research and development						100,000	700,000	200,000	2,700,000
Biogen Inc [Member] | Maximum [Member]
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Potential amount eligible to receive for certain milestones		$ 126,300,000
Potential amount to be funded for achievement of specified commercialized and sales milestones		$ 167,500,000
Shire AG [Member]
Research and Development Arrangement, Contract to Perform for Others [Line Items]
Revenues under agreement					$ 13,000,000
Deferred revenue						2,900,000		2,900,000
Research and development						200,000	$ 3,200,000	$ 900,000	$ 11,900,000
Aggregate number of gene targets | Targets					7
Number of initial gene targets | Targets					4
Number of gene targets | Targets				5
Number of additional gene targets | Targets				2				1
Recognition of milestone										$ 1,000,000
Amount reimbursed and recognized related to prior obligations	$ 3,400,000					$ 0		$ 0
Research program to develop laboratory research reagents								6 years

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