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Stock-Based Compensation (Tables)
 9 Months Ended
Sep. 30, 2017
Disclosure Of Compensation Related Costs Sharebased Payments [Abstract]  
Stock-Based Compensation Expense

The following table shows total stock-based compensation expense included in the condensed consolidated statements of operations for the three and nine months ended September 30, 2017 and 2016 (in thousands):

 

 

 

Three Months Ended

 

 

Nine Months Ended

 

 

 

September 30,

 

 

September 30,

 

 

 

2017

 

 

2016

 

 

2017

 

 

2016

 

Research and development

 

$

1,331

 

 

$

1,790

 

 

$

3,766

 

 

$

5,368

 

General and administrative

 

 

888

 

 

 

945

 

 

 

3,203

 

 

 

7,760

 

Total stock-based compensation expense

 

$

2,219

 

 

$

2,735

 

 

$

6,969

 

 

$

13,128