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Major Customers, Partnerships and Strategic Alliances - Agreement with Dow AgroSciences in Plant Agriculture - Additional Information (Detail) - Dow Agro Sciences [Member] - License agreement terms [Member] - USD ($)
12 Months Ended
Dec. 31, 2017
Dec. 31, 2016
Dec. 31, 2015
Dec. 31, 2005
Research and Development Arrangement, Contract to Perform for Others [Line Items]        
Research program to develop laboratory research reagents       3 years
One-time license fee earned on exercise of option       $ 6,000,000
Royalty revenues       $ 2,300,000
Percentage of royalties to be received from sublicensing       25.00%
Previously agreed research, development and commercialization milestone payments, and royalties on sales of products       $ 4,000,000
Fee due       $ 25,300,000
Minimum license annual fees specific reckoning period       11 years
Revenue attributable to research and development $ 3,000,000 $ 5,100,000 $ 3,000,000  
Minimum [Member]        
Research and Development Arrangement, Contract to Perform for Others [Line Items]        
Annual fees       $ 250,000
Maximum [Member]        
Research and Development Arrangement, Contract to Perform for Others [Line Items]        
Annual fees       $ 3,000,000