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Major Customers, Partnerships and Strategic Alliances - Additional Information (Detail) |  | 1 Months Ended | 3 Months Ended | 6 Months Ended | 12 Months Ended
--- | --- | --- | --- | --- | ---
Apr. 05, 2018 USD ($) | May 31, 2018 USD ($) | Apr. 30, 2018 USD ($) | Feb. 28, 2018 | Dec. 31, 2017 USD ($) | May 31, 2017 USD ($) | Jan. 31, 2014 USD ($) Program | Jan. 31, 2012 USD ($) | Jun. 30, 2018 USD ($) | Jun. 30, 2017 USD ($) | Jun. 30, 2018 USD ($) Product Milestone | Jun. 30, 2017 USD ($) | Dec. 31, 2014 USD ($)
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | $ 21,416,000 | $ 8,253,000 | $ 34,053,000 | $ 11,678,000 |
Number of research programs | Program |  |  |  |  |  |  | 2 |  |  |  |  |  |
Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | 21,289,000 | 7,977,000 | 33,840,000 | 11,283,000 |
Revenue from Grants [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | 127,000 | 276,000 | 213,000 | 395,000 |
California Institute for Regenerative Medicine ("CIRM") [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  | 1,700,000 |  | 1,700,000 |  |
Funds due under the agreement |  | $ 8,000,000 |  |  |  |  |  |  |  |  |  |  |
California Institute for Regenerative Medicine ("CIRM") [Member] | Revenue from Grants [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | $ 0 |  | $ 0 |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Collaboration waiting period under HSR expiration date at which time agreement became effective |  |  |  | Apr. 05, 2018 |  |  |  |  |  |  |  |  |
Milestone payments received |  |  | $ 150,000,000 |  |  |  |  |  |  |  |  |  |
Initial research term of agreement |  |  |  |  |  |  |  |  |  |  | 6 years |  |
Research agreement term |  |  |  |  |  |  |  |  |  |  | an option to extend the research term of the agreement for up to two additional one-year periods for a separate upfront fee of $10.0 million per year. |  |
Lease agreement, extendable lease term |  |  |  |  |  |  |  |  | 2 years |  | 2 years |  |
Separate upfront fee |  |  |  |  |  |  |  |  |  |  | $ 10,000,000 |  |
Deferred revenue |  |  |  |  |  |  |  |  | $ 144,000,000 |  | 144,000,000 |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Upfront License Fee [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  | 185,900,000 |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | $ 3,010,000,000 |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Achievement of Specified Research, Clinical Development, Regulatory and First Commercial Sale Milestones [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | 1,260,000,000 |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Achievement of Specified Sales-based Milestones if Annual Worldwide Net Sales of Licensed Products Reach Specified Levels [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | $ 1,750,000,000 |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Research Services [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | 1,500,000 |  |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | 6,000,000 |  |  |  |
Pfizer [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  | 0 |  |
Revenues under agreement |  |  |  |  |  | $ 70,000,000 |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  | 31,900,000 |  | $ 31,900,000 |  |
Number of products approved | Product |  |  |  |  |  |  |  |  |  |  | 0 |  |
Pfizer [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount eligible to receive for certain milestones |  |  |  |  |  | 208,500,000 |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  | 475,000,000 |  |  |  |  |  |  |
Pfizer [Member] | Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | 8,100,000 |  | $ 15,800,000 |  |
Pfizer [Member] | SB-525 [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  |  | 0 |  |
Pfizer [Member] | SB-525 [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  |  | 266,500,000 |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  | 300,000,000 |  |  |  |  |  |  |
Pfizer [Member] | Other Products [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  | $ 175,000,000 |  |  |  |  |  |  |
Pfizer [Member] | Royalty Revenues [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  | $ 0 |  |
Pfizer [Member] | C9ORF72 [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  | $ 12,000,000 |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  | 10,900,000 |  | 10,900,000 |  |
Revenues |  |  |  |  |  |  |  |  | 600,000 |  | $ 1,100,000 |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  |  | 0 |  |
Pfizer [Member] | C9ORF72 [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount eligible to receive for certain milestones |  |  |  |  | 60,000,000 |  |  |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  | 90,000,000 |  |  |  |  |  |  |  |
Bioverativ Inc [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  | $ 0 |  |
Revenues under agreement |  |  |  |  |  |  | $ 20,000,000 |  |  |  | 75,700,000 |  |
Deferred revenue |  |  |  |  |  |  |  |  | 6,200,000 |  | 6,200,000 |  |
Revenues |  |  |  |  |  |  |  |  | 4,516,000 | 3,510,000 | $ 8,898,000 | 5,661,000 |
Number of products approved | Product |  |  |  |  |  |  |  |  |  |  | 0 |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  |  | 0 |  |
Bioverativ Inc [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount eligible to receive for certain milestones |  |  |  |  |  |  | 115,800,000 |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  |  |  | $ 160,500,000 |  |  |  |  |  |
Bioverativ Inc [Member] | Research Services [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | 3,332,000 | 3,068,000 | $ 6,560,000 | 4,777,000 |
Bioverativ Inc [Member] | Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | $ 1,184,000 | 442,000 | 2,338,000 | 884,000 |
Bioverativ Inc [Member] | Royalty Revenues [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  | $ 0 |  |
Shire AG [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  | $ 13,000,000 |  |  |  |  |
Deferred revenue |  |  |  |  | $ 2,300,000 |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  | 586,000 |  | 1,277,000 |
Recognition of milestone |  |  |  |  |  |  |  |  |  |  |  |  | $ 1,000,000
Shire AG [Member] | Research Services [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  | 3,000 |  | 110,000 |
Shire AG [Member] | Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  | $ 583,000 |  | $ 1,167,000 |

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