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Organization, Basis of Presentation and Summary of Significant Accounting Policies (Tables) | 9 Months Ended
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Sep. 30, 2018
Summary of Reconciliation of Cash, Cash Equivalents and Restricted Cash | The following table provides a reconciliation of cash, cash equivalents, and restricted cash within the statement of financial position that sum to the total of the same amounts in the statement of cash flows:
|  | As of September 30, |
(in thousands) |  | 2018 |  |  | 2017 |
Cash and cash equivalents |  | $ | 39,298 |  |  | $ | 29,894 |
Restricted cash included in other non-current assets |  |  | 79,941 |  |  |  | — |
Total cash, cash equivalents, and restricted cash shown in the statements of cash flows |  | $ | 119,239 |  |  | $ | 29,894 |
Topic 606 [Member] |
Summary of Financial Statement Line Item Affected Cumulative Adjustment and Compared with Guidance in Effect Prior to Adoption | Refer below for a summary of the amount by which each financial statement line item was affected by the impact of the cumulative adjustment and as compared with the guidance that was in effect prior to the adoption:
|  | Impact of Topic 606 Adoption on Condensed Consolidated Balance Sheet as of January 1, 2018 |
(in thousands) |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |
Deferred revenue, current portion |  | $ | 29,626 |  |  | $ | 1,281 |  |  | $ | 28,345 |
Deferred revenue, noncurrent portion |  | $ | 26,846 |  |  | $ | (2,398 | ) |  | $ | 29,244 |
Accumulated deficit |  | $ | (494,362 | ) |  | $ | 1,117 |  |  | $ | (495,479 | )
|  | Impact of Topic 606 Adoption on Condensed Consolidated Balance Sheet as of September 30, 2018 |
(in thousands) |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |
Deferred revenue, current portion |  | $ | 51,094 |  |  | $ | 12,741 |  |  | $ | 63,835 |
Deferred revenue, noncurrent portion |  | $ | 123,917 |  |  | $ | (3,130 | ) |  | $ | 120,787 |
Accumulated deficit |  | $ | (544,032 | ) |  | $ | (9,611 | ) |  | $ | (553,643 | )
|  | Impact of Topic 606 Adoption on Condensed Consolidated Statement of Operations and Comprehensive Loss for the Three Months Ended September 30, 2018 |  |  | Impact of Topic 606 Adoption on Condensed Consolidated Statement of Operations and Comprehensive Loss for the Nine Months Ended September 30, 2018 |
(in thousands) |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |  |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |
Collaboration revenue |  | $ | 23,538 |  |  | $ | (4,461 | ) |  | $ | 19,077 |  |  | $ | 57,378 |  |  | $ | (8,495 | ) |  | $ | 48,883 |
Net loss |  | $ | (12,843 | ) |  | $ | (4,461 | ) |  | $ | (17,304 | ) |  | $ | (49,670 | ) |  | $ | (8,495 | ) |  | $ | (58,165 | )
Net loss per share - basic and diluted: |  | $ | (0.13 | ) |  | $ | (0.04 | ) |  | $ | (0.17 | ) |  | $ | (0.52 | ) |  | $ | (0.09 | ) |  | $ | (0.61 | )
|  | Impact of Topic 606 Adoption on Condensed Consolidated Statement of Cash Flows for the Nine Months Ended September 30, 2018 |
(in thousands) |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |
Net loss |  | $ | (49,670 | ) |  | $ | (8,495 | ) |  | $ | (58,165 | )
Changes in deferred revenue |  | $ | 118,540 |  |  | $ | 8,495 |  |  | $ | 127,035 |

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