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Major Customers, Partnerships and Strategic Alliances - Additional Information (Detail) |  | 1 Months Ended | 3 Months Ended | 12 Months Ended
--- | --- | --- | --- | ---
Apr. 05, 2018 USD ($) | May 31, 2018 USD ($) | Apr. 30, 2018 USD ($) | Feb. 28, 2018 | Dec. 31, 2017 USD ($) | May 31, 2017 USD ($) | Jan. 31, 2014 USD ($) Program | Jan. 31, 2012 USD ($) | Dec. 31, 2018 USD ($) | Sep. 30, 2018 USD ($) | Jun. 30, 2018 USD ($) | Mar. 31, 2018 USD ($) | Dec. 31, 2017 USD ($) | Sep. 30, 2017 USD ($) | Jun. 30, 2017 USD ($) | Mar. 31, 2017 USD ($) | Dec. 31, 2018 USD ($) Product Milestone | Dec. 31, 2017 USD ($) | Dec. 31, 2016 USD ($) | Dec. 31, 2014 USD ($)
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | $ 26,837,000 | $ 23,562,000 | $ 21,416,000 | $ 12,637,000 | $ 13,077,000 | $ 11,812,000 | $ 8,253,000 | $ 3,425,000 | $ 84,452,000 | $ 36,567,000 | $ 19,389,000 |
Number of research programs | Program |  |  |  |  |  |  | 2 |  |  |  |  |  |  |  |  |  |  |  |  |
Other long-term liabilities |  |  |  |  |  |  |  |  | 1,960,000 |  |  |  |  |  |  |  | 1,960,000 |  |  |
Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 84,065,000 | 35,960,000 | 18,881,000 |
Research Grants [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 387,000 | 607,000 | 508,000 |
California Institute for Regenerative Medicine ("CIRM") [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Funds due under the agreement |  | $ 8,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Other long-term liabilities |  |  |  |  |  |  |  |  | 1,800,000 |  |  |  |  |  |  |  | 1,800,000 |  |  |
Interest payable |  |  |  |  |  |  |  |  | $ 100,000 |  |  |  |  |  |  |  | 100,000 |  |  |
California Institute for Regenerative Medicine ("CIRM") [Member] | Research Grants [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 1,700,000 |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaboration waiting period under HSR expiration date at which time agreement became effective |  |  |  | Apr. 05, 2018 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  | $ 150,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Initial research term of agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 6 years |  |  |
Research agreement term |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | an option to extend the research term for up to two additional one-year periods for a separate fee of $10.0 million per year. |  |  |
Lease agreement, extendable lease term |  |  |  |  |  |  |  |  | 2 years |  |  |  |  |  |  |  | 2 years |  |  |
Separate fee for additional term |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 10,000,000 |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 185,900,000 |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 150,000,000 |  |  |
Collaborative arrangement estimated reimbursable service costs |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 35,900,000 |  |  |
Deferred revenue |  |  |  |  |  |  |  |  | $ 131,500,000 |  |  |  |  |  |  |  | 131,500,000 |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 18,500,000 |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Research Services [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,000,000 |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received | $ 3,010,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Achievement of Specified Research, Clinical Development, Regulatory and First Commercial Sale Milestones [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | 1,260,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma, Inc. ("Kite") [Member] | Collaboration and License Agreement [Member] | Achievement of Specified Sales-based Milestones if Annual Worldwide Net Sales of Licensed Products Reach Specified Levels [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | $ 1,750,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |
Revenues under agreement |  |  |  |  |  | $ 70,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  | 10,000,000 |  |  |  |  |  |  |  | $ 10,000,000 |  |  |
Number of products approved | Product |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |
Pfizer [Member] | Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 37,800,000 | 17,000,000 |  |
Pfizer [Member] | SB-525 [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |
Pfizer [Member] | Royalty Revenues [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 0 |  |  |
Pfizer [Member] | C9ORF72 [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  | $ 12,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  | 9,800,000 |  |  |  |  |  |  |  | 9,800,000 |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 2,200,000 |  |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |
Pfizer [Member] | Maximum [Member] | SB-525 [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  |  | 266,500,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  | 300,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer [Member] | Maximum [Member] | Other Products [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  | 175,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer [Member] | Achievement of Specified Clinical Development Intellectual Property and Regulatory Milestones | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  | 208,500,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer [Member] | Achievement of First Commercial Sale Milestones | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  | $ 475,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer [Member] | Achievement of Specified Preclinical Development Clinical Development and First Commercial Sale Milestones | Maximum [Member] | C9ORF72 [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  | 60,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer [Member] | Achievement of Commercial Milestones | Maximum [Member] | C9ORF72 [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  | 90,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Bioverativ Inc [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 0 |  |  |
Revenues under agreement |  |  |  |  |  |  | $ 20,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 13,516,000 | 12,258,000 | 8,886,000 |
Number of products approved | Product |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |
Bioverativ Inc [Member] | Collaboration Agreements [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 4,013,000 | 1,769,000 | 2,321,000 |
Bioverativ Inc [Member] | Research Services [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 9,503,000 | 10,489,000 | 6,565,000 |
Bioverativ Inc [Member] | Collaboration and License Agreement [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 75,700,000 |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 20,000,000 |  |  |
Collaborative arrangement estimated reimbursable service costs |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 55,700,000 |  |  |
Deferred revenue |  |  |  |  |  |  |  |  | $ 4,600,000 |  |  |  |  |  |  |  | $ 4,600,000 |  |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |
Bioverativ Inc [Member] | Collaboration and License Agreement [Member] | Royalty Revenues [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 0 |  |  |
Bioverativ Inc [Member] | Collaboration and License Agreement [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  | 276,300,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Bioverativ Inc [Member] | Collaboration and License Agreement [Member] | Achievement Of Specified Clinical Development And Regulatory Milestones [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  | 115,800,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Bioverativ Inc [Member] | Collaboration and License Agreement [Member] | Achievement of Specified Sales Milestones [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  | $ 160,500,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Shire AG [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 0 | 2,400,000 | $ 3,300,000 |
Shire AG [Member] | Collaboration and License Agreement [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  | $ 13,000,000 |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  | $ 2,300,000 |  |  |  |  |  |  |  | $ 2,300,000 |  |  |  |  | $ 2,300,000 |  |
Recognition of milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 1,000,000

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