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Organization and Summary of Significant Accounting Policies (Tables) | 12 Months Ended
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Dec. 31, 2018
Schedule of Goodwill | During the fourth quarter of 2018, the Company performed an assessment of the qualitative factors affecting the fair value of its reporting unit and concluded that it was not more likely than not that the fair value of its reporting unit was less than carrying value and that, as a result, it is not more likely than not that goodwill is impaired.
Balance as of December 31, 2017 |  | $ | 1,585 |
Goodwill acquired |  |  | 38,995 |
Foreign currency translations adjustment |  |  | (536 | )
Balance as of December 31, 2018 |  | $ | 40,044 |
Summary of Reconciliation of Cash, Cash Equivalents and Restricted Cash | The following table provides a reconciliation of cash, cash equivalents, and restricted cash within the consolidated statements of cash flows that sum to the total of the same amounts in the statement of cash flows for the years ended December 31, 2018 and 2017, respectively (in thousands):
|  | Year Ended December 31, |
|  | 2018 |  |  | 2017 |
Cash and cash equivalents |  | $ | 140,418 |  |  | $ | 49,826 |
Restricted cash |  |  | 3,500 |  |  |  | 3,500 |
Total cash, cash equivalents, and restricted cash |  | $ | 143,918 |  |  | $ | 53,326 |
Topic 606 [Member] |
Summary of Financial Statement Line Item Affected Cumulative Adjustment and Compared with Guidance in Effect Prior to Adoption | Refer below for a summary of the amount by which each financial statement line item that was affected by the impact of the cumulative adjustment and as compared with the guidance that was in effect prior to the adoption:
|  | Impact of Topic 606 Adoption on Consolidated Balance Sheet as of January 1, 2018 |
(in thousands) |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |
Deferred revenue, current portion |  | $ | 29,626 |  |  | $ | 1,281 |  |  | $ | 28,345 |
Deferred revenue, noncurrent portion |  |  | 26,846 |  |  |  | (2,398 | ) |  |  | 29,244 |
Accumulated deficit |  |  | (494,362 | ) |  |  | 1,117 |  |  |  | (495,479 | )
|  | Impact of Topic 606 Adoption on Consolidated Balance Sheet as of December 31, 2018 |
(in thousands) |  | As reported under Topic 606 |  |  |  |  | Adjustments |  |  |  |  | Balances without adoption of Topic 606 |
Deferred revenue, current portion |  | $ | 47,564 |  |  |  |  | $ | 15,553 |  |  |  |  | $ | 63,117 |
Deferred revenue, noncurrent portion |  |  | 108,273 |  |  |  |  |  | (879 | ) |  |  |  |  | 107,394 |
Accumulated deficit |  |  | (562,696 | ) |  |  |  |  | (14,674 | ) |  |  |  |  | (577,370 | )
|  | Impact of Topic 606 Adoption on Consolidated Statement of Operations and Comprehensive Loss for the Year Ended December 31, 2018 |
(in thousands, except per share amounts) |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |
Collaboration revenue |  | $ | 84,065 |  |  | $ | (13,558 | ) |  | $ | 70,507 |
Net loss |  |  | (68,334 | ) |  |  | (13,558 | ) |  |  | (81,892 | )
Net loss per share - basic and diluted: |  |  | (0.70 | ) |  |  | (0.14 | ) |  |  | (0.84 | )
|  | Impact of Topic 606 Adoption on Consolidated Statement of Cash Flows for the Year Ended December 31, 2018 |
(in thousands) |  | As reported under Topic 606 |  |  | Adjustments |  |  | Balances without adoption of Topic 606 |
Net loss |  | $ | (68,334 | ) |  | $ | (13,558 | ) |  | $ | (81,892 | )
Changes in deferred revenue |  |  | 99,364 |  |  |  | 13,558 |  |  |  | 112,922 |

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