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Major Customers, Partnerships and Strategic Alliances - Agreement with Dow AgroSciences in Plant Agriculture - Additional Information (Detail) - USD ($) | 3 Months Ended | 12 Months Ended
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Dec. 31, 2018 | Sep. 30, 2018 | Jun. 30, 2018 | Mar. 31, 2018 | Dec. 31, 2017 | Sep. 30, 2017 | Jun. 30, 2017 | Mar. 31, 2017 | Dec. 31, 2018 | Dec. 31, 2017 | Dec. 31, 2016 | Dec. 31, 2005
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues | $ 26,837,000 | $ 23,562,000 | $ 21,416,000 | $ 12,637,000 | $ 13,077,000 | $ 11,812,000 | $ 8,253,000 | $ 3,425,000 | $ 84,452,000 | $ 36,567,000 | $ 19,389,000 |
Dow Agro Sciences [Member] | License agreement terms [Member] |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Research program to develop laboratory research reagents |  |  |  |  |  |  |  |  |  |  |  | 3 years
One-time license fee earned on exercise of option |  |  |  |  |  |  |  |  |  |  |  | $ 6,000,000
Percentage of royalties to be received from sublicensing |  |  |  |  |  |  |  |  |  |  |  | 25.00%
Previously agreed research, development and commercialization milestone payments, and royalties on sales of products |  |  |  |  |  |  |  |  |  |  |  | $ 4,000,000
Fee due |  |  |  |  |  |  |  |  |  |  |  | $ 25,300,000
Minimum license annual fees specific reckoning period |  |  |  |  |  |  |  |  |  |  |  | 11 years
Dow Agro Sciences [Member] | License agreement terms [Member] | Minimum [Member] |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Annual fees |  |  |  |  |  |  |  |  |  |  |  | $ 250,000
Dow Agro Sciences [Member] | License agreement terms [Member] | Maximum [Member] |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Annual fees |  |  |  |  |  |  |  |  |  |  |  | 3,000,000
Dow Agro Sciences [Member] | License agreement terms [Member] | Royalty Revenues [Member] |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | $ 2,300,000
Dow Agro Sciences [Member] | License agreement terms [Member] | License Fee Revenues [Member] |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  | $ 3,000,000 | $ 3,000,000 | $ 5,100,000 |

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