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Condensed Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-in Capital | Accumulated Deficit | Accumulated Other Comprehensive Income (Loss) | Non- Controlling Interest
--- | --- | --- | --- | --- | --- | ---
Beginning Balances (in shares) at Dec. 31, 2017 |  | 85,598 |  |  |  |
Beginning Balances at Dec. 31, 2017 | $ 187,900 | $ 856 | $ 682,809 | $ (495,479) | $ (286) |
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |
Cumulative-effect adjustment of ASC | Topic 606 | 1,117 |  |  | 1,117 |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  | 1,443 |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 10,584 | $ 14 | 10,570 |  |  |
Stock-based compensation | 3,050 |  | 3,050 |  |  |
Net unrealized gain (loss) on marketable securities | (99) |  |  |  | (99) |
Net loss | (20,187) |  |  | (20,187) |  |
Ending Balances (in shares) at Mar. 31, 2018 |  | 87,041 |  |  |  |
Ending Balances at Mar. 31, 2018 | 182,365 | $ 870 | 696,429 | (514,549) | (385) |
Beginning Balances (in shares) at Dec. 31, 2017 |  | 85,598 |  |  |  |
Beginning Balances at Dec. 31, 2017 | 187,900 | $ 856 | 682,809 | (495,479) | (286) |
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |
Net unrealized gain (loss) on marketable securities | 131 |  |  |  |  |
Net loss | (36,827) |  |  |  |  |
Ending Balances (in shares) at Jun. 30, 2018 |  | 101,624 |  |  |  |
Ending Balances at Jun. 30, 2018 | 387,867 | $ 1,016 | 918,197 | (531,189) | (157) |
Beginning Balances (in shares) at Mar. 31, 2018 |  | 87,041 |  |  |  |
Beginning Balances at Mar. 31, 2018 | 182,365 | $ 870 | 696,429 | (514,549) | (385) |
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  | 263 |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 1,954 | $ 2 | 1,952 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  | 163 |  |  |  |
Issuance of common stock under employee stock purchase plan | 690 | $ 2 | 688 |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  | 14,157 |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 215,756 | $ 142 | 215,614 |  |  |
Stock-based compensation | 3,514 |  | 3,514 |  |  |
Foreign currency translation adjustment | (2) |  |  |  | (2) |
Net unrealized gain (loss) on marketable securities | 230 |  |  |  | 230 |
Net loss | (16,640) |  |  | (16,640) |  |
Ending Balances (in shares) at Jun. 30, 2018 |  | 101,624 |  |  |  |
Ending Balances at Jun. 30, 2018 | 387,867 | $ 1,016 | 918,197 | (531,189) | (157) |
Beginning Balances (in shares) at Dec. 31, 2018 |  | 102,188 |  |  |  |
Beginning Balances at Dec. 31, 2018 | 367,257 | $ 1,022 | 929,632 | (562,696) | (1,440) | $ 739
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |
Cumulative-effect adjustment of ASC | Topic 842 | 897 |  |  | 897 |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  | 141 |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 216 | $ 1 | 215 |  |  |
Issuance costs related to public offering and TxCell Acquisition | (258) |  | (258) |  |  |
Stock-based compensation | 4,523 |  | 4,523 |  |  |
Foreign currency translation adjustment | (1,504) |  |  |  | (1,504) |
Net unrealized gain (loss) on marketable securities | 253 |  |  |  | 253 |
Net loss | (42,203) |  |  | (42,150) |  | (53)
Ending Balances (in shares) at Mar. 31, 2019 |  | 102,329 |  |  |  |
Ending Balances at Mar. 31, 2019 | 329,181 | $ 1,023 | 934,112 | (603,949) | (2,691) | 686
Beginning Balances (in shares) at Dec. 31, 2018 |  | 102,188 |  |  |  |
Beginning Balances at Dec. 31, 2018 | 367,257 | $ 1,022 | 929,632 | (562,696) | (1,440) | 739
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |
Net unrealized gain (loss) on marketable securities | 737 |  |  |  |  |
Net loss | (72,559) |  |  |  |  |
Ending Balances (in shares) at Jun. 30, 2019 |  | 115,603 |  |  |  |
Ending Balances at Jun. 30, 2019 | 445,748 | $ 1,156 | 1,078,976 | (634,233) | (765) | 614
Beginning Balances (in shares) at Mar. 31, 2019 |  | 102,329 |  |  |  |
Beginning Balances at Mar. 31, 2019 | 329,181 | $ 1,023 | 934,112 | (603,949) | (2,691) | 686
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  | 492 |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 2,444 | $ 5 | 2,439 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  | 132 |  |  |  |
Issuance of common stock under employee stock purchase plan | 1,138 | $ 1 | 1,137 |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  | 12,650 |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 136,566 | $ 127 | 136,439 |  |  |
Issuance costs related to public offering and TxCell Acquisition | (18) |  | (18) |  |  |
Stock-based compensation | 4,867 |  | 4,867 |  |  |
Foreign currency translation adjustment | 1,442 |  |  |  | 1,442 |
Net unrealized gain (loss) on marketable securities | 484 |  |  |  | 484 |
Net loss | (30,356) |  |  | (30,284) |  | (72)
Ending Balances (in shares) at Jun. 30, 2019 |  | 115,603 |  |  |  |
Ending Balances at Jun. 30, 2019 | $ 445,748 | $ 1,156 | $ 1,078,976 | $ (634,233) | $ (765) | $ 614

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