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Major Customers, Partnerships and Strategic Alliances - Additional Information (Detail) |  | 1 Months Ended | 3 Months Ended | 6 Months Ended | 7 Months Ended | 13 Months Ended | 25 Months Ended | 65 Months Ended
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Apr. 05, 2018 USD ($) | May 31, 2018 USD ($) | Apr. 30, 2018 USD ($) | Dec. 31, 2017 USD ($) milestone | May 31, 2017 USD ($) | Jan. 31, 2014 USD ($) program | Jun. 30, 2019 USD ($) | Mar. 31, 2019 USD ($) | Jun. 30, 2018 USD ($) | Jun. 30, 2019 USD ($) Milestone | Jun. 30, 2018 USD ($) | Dec. 31, 2018 USD ($) | Jun. 30, 2019 USD ($) | Jun. 30, 2019 USD ($) Product | Jun. 30, 2019 USD ($) Product
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  | $ 17,548,000 |  | $ 21,416,000 | $ 25,619,000 | $ 34,053,000 |  |  |  |
Number of research programs | program |  |  |  |  |  | 2 |  |  |  |  |  |  |  |  |
Other long-term liabilities |  |  |  |  |  |  | 3,288,000 |  |  | 3,288,000 |  | $ 1,960,000 | $ 3,288,000 | $ 3,288,000 | $ 3,288,000
California Institute For Regenerative Medicine Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Funds due under the agreement |  | $ 8,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |
Other long-term liabilities |  |  |  |  |  |  | 3,300,000 |  |  | 3,300,000 |  | 1,800,000 | 3,300,000 | 3,300,000 | 3,300,000
California Institute For Regenerative Medicine Agreement | Research Grants |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | 1,700,000 | 3,000,000.0 |  |
Kite Pharma Inc |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  | 9,060,000 |  | 7,515,000 | 17,372,000 | 7,515,000 |  |  |  |
Kite Pharma Inc | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  | $ 6,227,000 |  | 5,953,000 | 12,386,000 | 5,953,000 |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaboration waiting period under HSR expiration date at which time agreement became effective | Apr. 05, 2018 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  | $ 150,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |
Initial research term of agreement | 6 years |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lease agreement, extendable lease term | 2 years |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research agreement term |  |  |  |  |  |  | an option to extend the research term of the agreement for up to two additional one-year periods for a separate upfront fee of $10.0 million per year. |  |  |  |  |  |  |  |
Separate upfront fee | $ 10,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price | 185,900,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement | 150,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs | 35,900,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  | $ 119,100,000 |  |  | 119,100,000 |  | 131,500,000 | 119,100,000 | 119,100,000 | 119,100,000
Kite Pharma Inc | Collaboration and License Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received | 3,010,000,000.00 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement | Achievement of Specified Research, Clinical Development, Regulatory and First Commercial Sale Milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | 1,260,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement | Achievement of Specified Sales-based Milestones if Annual Worldwide Net Sales of Licensed Products Reach Specified Levels |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | $ 1,750,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  | $ 12,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  | $ 70,000,000.0 |  |  |  |  |  |  |  |  | 0 |
Deferred revenue |  |  |  |  |  |  | 14,300,000 |  |  | 14,300,000 |  | 10,000,000.0 | 14,300,000 | $ 14,300,000 | 14,300,000
Number of products approved | Product |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |
Revenues |  |  |  |  |  |  |  |  |  | 25,600,000 |  |  |  |  |
Pfizer | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenue reduction due to decrease in measure of proportional cumulative performance |  |  |  |  |  |  |  | $ 3,000,000.0 |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  | 455,000 |  | 617,000 | $ 1,070,000 | 1,076,000 |  |  |  |
Pfizer | SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  | 0 |  |  |  |  |
Pfizer | Royalty Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 0 |
Pfizer | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  | 8,700,000 |  |  | $ 8,700,000 |  | 9,800,000 | 8,700,000 | 8,700,000 | 8,700,000
Number of milestones included in transaction price | milestone |  |  |  | 0 |  |  |  |  |  |  |  |  |  |  |
Pfizer | Maximum | SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  | 266,500,000 |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  | 300,000,000.0 |  |  |  |  |  |  |  |  |  |
Pfizer | Maximum | Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  | 175,000,000.0 |  |  |  |  |  |  |  |  |  |
Pfizer | Achievement of Specified Clinical Development Intellectual Property and Regulatory Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  | 208,500,000 |  |  |  |  |  |  |  |  |  |
Pfizer | Achievement of First Commercial Sale Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  | $ 475,000,000.0 |  |  |  |  |  |  |  |  |  |
Pfizer | Achievement of Specified Preclinical Development Clinical Development and First Commercial Sale Milestones | Maximum | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  | $ 60,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Pfizer | Achievement of Commercial Milestones | Maximum | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  | $ 90,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Pfizer SB-525 | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  | 4,635,000 |  | 8,183,000 | 1,595,000 | 15,841,000 |  |  |  |
Bioverativ Inc |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  | 3,059,000 |  | 4,516,000 | 5,009,000 | 8,898,000 |  |  |  |
Bioverativ Inc | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  | 901,000 |  | $ 1,184,000 | $ 1,654,000 | $ 2,338,000 |  |  |  |
Bioverativ Inc | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  | $ 75,700,000 |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  | 20,000,000.0 |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs |  |  |  |  |  | 55,700,000 |  |  |  |  |  |  |  |  |
Number of milestones included in transaction price | Milestone |  |  |  |  |  |  |  |  |  | 0 |  |  |  |  |
Bioverativ Inc | Collaboration and License Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  | 276,300,000 |  |  |  |  |  |  |  |  |
Sanofi |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 0
Revenues under agreement |  |  |  |  |  | 20,000,000.0 |  |  |  |  |  |  |  |  |
Number of products approved | Product |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0
Sanofi | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  | $ 2,900,000 |  |  | $ 2,900,000 |  | $ 4,600,000 | $ 2,900,000 | $ 2,900,000 | $ 2,900,000
Sanofi | Collaboration and License Agreement | Royalty Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 0
Sanofi | Collaboration and License Agreement | Achievement Of Specified Clinical Development And Regulatory Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  | 115,800,000 |  |  |  |  |  |  |  |  |
Sanofi | Collaboration and License Agreement | Achievement of Specified Sales Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  | $ 160,500,000 |  |  |  |  |  |  |  |  |

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