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Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Common Stock | Additional Paid-in Capital | Accumulated Deficit | Accumulated Other Comprehensive Income/ (Loss) | Non- Controlling Interest
--- | --- | --- | --- | --- | --- | ---
Beginning Balances (in shares) at Dec. 31, 2016 |  | 70,871,902 |  |  |  |
Beginning Balances at Dec. 31, 2016 | $ 136,195 | $ 709 | $ 576,377 | $ (440,911) | $ 20 | $ 0
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  | 2,101,489 |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | $ 15,099 | $ 21 | 15,078 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) | 253,994 | 253,994 |  |  |  |
Issuance of common stock under employee stock purchase plan | $ 818 | $ 2 | 816 |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  | 12,371,149 |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 81,573 | $ 124 | 81,449 |  |  |
Stock-based compensation | 9,089 |  | 9,089 |  |  |
Net pension losses | 0 |  |  |  |  |
Net unrealized loss on marketable securities, net of tax | (306) |  |  |  | (306) |
Net loss | (54,568) |  |  | (54,568) |  |
Ending Balances (in shares) at Dec. 31, 2017 |  | 85,598,534 |  |  |  |
Ending Balances at Dec. 31, 2017 | 187,900 | $ 856 | 682,809 | (495,479) | (286) | 0
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  | 2,103,727 |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | $ 14,467 | $ 20 | 14,447 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) | 328,710 | 328,710 |  |  |  |
Issuance of common stock under employee stock purchase plan | $ 1,484 | $ 4 | 1,480 |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  | 14,156,500 |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 215,758 | $ 142 | 215,616 |  |  |
Stock-based compensation | 14,677 |  | 14,677 |  |  |
Acquisition of additional shares of Sangamo France | 603 |  | 603 |  |  |
Non-controlling interest upon Acquisition of Sangamo France | 1,313 |  |  |  |  | 1,313
Foreign currency translation adjustment | (1,148) |  |  |  | (1,129) | (19)
Net pension losses | (21) |  |  |  | (21) |
Net unrealized loss on marketable securities, net of tax | (4) |  |  |  | (4) |
Net loss | $ (68,889) |  |  | (68,334) |  | (555)
Ending Balances (in shares) at Dec. 31, 2018 | 102,187,471 | 102,187,471 |  |  |  |
Ending Balances at Dec. 31, 2018 | $ 367,257 | $ 1,022 | 929,632 | (562,696) | (1,440) | 739
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  | 885,873 |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | $ 3,677 | $ 9 | 3,668 |  |  |
Issuance of common stock under employee stock purchase plan (in shares) | 249,364 | 249,364 |  |  |  |
Issuance of common stock under employee stock purchase plan | $ 2,044 | $ 2 | 2,042 |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  | 12,650,000 |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 136,308 | $ 127 | 136,181 |  |  |
Stock-based compensation | 19,330 |  | 19,330 |  |  |
Acquisition of additional shares of Sangamo France | (321) |  |  |  |  | (321)
Issuance costs related to Sangamo France Acquisition | 25 |  | 25 |  |  |
Foreign currency translation adjustment | (1,573) |  |  |  | (1,573) |
Net pension losses | (28) |  |  |  | (28) |
Net unrealized loss on marketable securities, net of tax | 592 |  |  |  | 592 |
Net loss | $ (95,419) |  |  | (95,186) |  | (233)
Ending Balances (in shares) at Dec. 31, 2019 | 115,972,708 | 115,972,708 |  |  |  |
Ending Balances at Dec. 31, 2019 | $ 432,739 | $ 1,160 | $ 1,090,828 | $ (656,985) | $ (2,449) | $ 185

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