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Consolidated Statements of Stockholders' Equity - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Deficit
Accumulated Other Comprehensive Income/ (Loss)
Non- Controlling Interest
Beginning Balances (in shares) at Dec. 31, 2016   70,871,902        
Beginning Balances at Dec. 31, 2016 $ 136,195 $ 709 $ 576,377 $ (440,911) $ 20 $ 0
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   2,101,489        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax $ 15,099 $ 21 15,078      
Issuance of common stock under employee stock purchase plan (in shares) 253,994 253,994        
Issuance of common stock under employee stock purchase plan $ 818 $ 2 816      
Issuance of common stock under public offering, net of issuance costs (in shares)   12,371,149        
Issuance of common stock under public offering, net of issuance costs 81,573 $ 124 81,449      
Stock-based compensation 9,089   9,089      
Net pension losses 0          
Net unrealized loss on marketable securities, net of tax (306)       (306)  
Net loss (54,568)     (54,568)    
Ending Balances (in shares) at Dec. 31, 2017   85,598,534        
Ending Balances at Dec. 31, 2017 187,900 $ 856 682,809 (495,479) (286) 0
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   2,103,727        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax $ 14,467 $ 20 14,447      
Issuance of common stock under employee stock purchase plan (in shares) 328,710 328,710        
Issuance of common stock under employee stock purchase plan $ 1,484 $ 4 1,480      
Issuance of common stock under public offering, net of issuance costs (in shares)   14,156,500        
Issuance of common stock under public offering, net of issuance costs 215,758 $ 142 215,616      
Stock-based compensation 14,677   14,677      
Acquisition of additional shares of Sangamo France 603   603      
Non-controlling interest upon Acquisition of Sangamo France 1,313         1,313
Foreign currency translation adjustment (1,148)       (1,129) (19)
Net pension losses (21)       (21)  
Net unrealized loss on marketable securities, net of tax (4)       (4)  
Net loss $ (68,889)     (68,334)   (555)
Ending Balances (in shares) at Dec. 31, 2018 102,187,471 102,187,471        
Ending Balances at Dec. 31, 2018 $ 367,257 $ 1,022 929,632 (562,696) (1,440) 739
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   885,873        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax $ 3,677 $ 9 3,668      
Issuance of common stock under employee stock purchase plan (in shares) 249,364 249,364        
Issuance of common stock under employee stock purchase plan $ 2,044 $ 2 2,042      
Issuance of common stock under public offering, net of issuance costs (in shares)   12,650,000        
Issuance of common stock under public offering, net of issuance costs 136,308 $ 127 136,181      
Stock-based compensation 19,330   19,330      
Acquisition of additional shares of Sangamo France (321)         (321)
Issuance costs related to Sangamo France Acquisition 25   25      
Foreign currency translation adjustment (1,573)       (1,573)
Net pension losses (28)       (28)  
Net unrealized loss on marketable securities, net of tax 592       592  
Net loss $ (95,419)     (95,186)   (233)
Ending Balances (in shares) at Dec. 31, 2019 115,972,708 115,972,708        
Ending Balances at Dec. 31, 2019 $ 432,739 $ 1,160 $ 1,090,828 $ (656,985) $ (2,449) $ 185