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v3.20.1

MAJOR CUSTOMERS, PARTNERSHIPS AND STRATEGIC ALLIANCES - Additional Information (Detail) |  | 1 Months Ended | 3 Months Ended | 16 Months Ended | 34 Months Ended |
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Apr. 05, 2018 USD ($) | Apr. 30, 2020 USD ($) $ / shares shares | Apr. 30, 2020 USD ($) $ / shares | Mar. 31, 2020 USD ($) $ / shares | Feb. 29, 2020 USD ($) product_target | Dec. 31, 2019 USD ($) | Sep. 30, 2019 USD ($) | May 31, 2018 USD ($) | Dec. 31, 2017 USD ($) milestone | May 31, 2017 USD ($) | Jan. 31, 2014 USD ($) program | Mar. 31, 2020 USD ($) option milestone product | Mar. 31, 2019 USD ($) | Sep. 30, 2019 USD ($) | Mar. 31, 2020 USD ($) | Feb. 28, 2020
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenue reduction due to decrease in measure of proportional cumulative performance, percentage |  |  |  |  |  |  |  |  |  |  |  |  | 38.00% |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | $ 13,076,000 | $ 8,071,000 |  |  |
Other long-term liabilities |  |  |  | $ 5,880,000 |  | $ 5,711,000 |  |  |  |  |  | 5,880,000 |  |  | $ 5,880,000 |
California Institute For Regenerative Medicine Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Funds due under the agreement |  |  |  |  |  |  |  | $ 8,000,000.0 |  |  |  |  |  |  |  |
Other long-term liabilities |  |  |  | 5,900,000 |  | 5,700,000 |  |  |  |  |  | 5,900,000 |  |  | 5,900,000 |
California Institute For Regenerative Medicine Agreement | Research Grants |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 5,200,000 |  |
Biogen MA, Inc. | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Proceeds from collaborators |  |  |  |  | $ 125,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  | $ 2,370,000,000 |  |  |  |  |  |  |  |  |  |  |
Number of product targets | product_target |  |  |  |  | 12 |  |  |  |  |  |  |  |  |  |  |
Number of product targets selected | product_target |  |  |  |  | 3 |  |  |  |  |  |  |  |  |  |  |
Number of additional product targets | product_target |  |  |  |  | 9 |  |  |  |  |  |  |  |  |  |  |
Target selection period |  |  |  |  | 5 years |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Collaboration and License Agreement | Subsequent Event |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Proceeds from collaborators |  |  | $ 125,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Collaboration and License Agreement | Pre-approval Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  | $ 925,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Collaboration and License Agreement | Sales-based Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  | $ 1,450,000,000 |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Stock Purchase Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Standstill restriction period |  |  |  |  | 3 years |  |  |  |  |  |  |  |  |  |  |
Standstill restriction, ownership threshold percentage ownership percentage |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5.00%
Agreement restriction, percentage of shares held |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 50.00%
Voting provisions expiration period |  |  |  |  | 2 years |  |  |  |  |  |  |  |  |  |  |
Voting provisions, ownership threshold percentage |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5.00%
Biogen MA, Inc. | Stock Purchase Agreement | Subsequent Event |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of shares issued in transaction (in shares) | shares |  | 24,420,157 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Sale of stock, price per share (in dollars per share) | $ / shares |  | $ 9.2137 | $ 9.2137 |  |  |  |  |  |  |  |  |  |  |  |  |
Consideration received on transaction |  | $ 225,000,000.0 | $ 225,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | 7,219,000 | 8,312,000 |  |  |
Kite Pharma Inc | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | $ 6,227,000 | 6,159,000 |  |  |
Kite Pharma Inc | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received | $ 150,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Initial research term of agreement | 6 years |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of options to extend initial research term | option |  |  |  |  |  |  |  |  |  |  |  | 2 |  |  |  |
Extended research term of agreement |  |  |  |  |  |  |  |  |  |  |  | 1 year |  |  |  |
Separate upfront fee | $ 10,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs for new research plan |  |  |  |  |  |  | $ 3,400,000 |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price | 189,300,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement | 150,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs | 39,300,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 100,300,000 |  | 106,500,000 |  |  |  |  |  | $ 100,300,000 |  |  | 100,300,000 |
Kite Pharma Inc | Collaboration and License Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received | 3,010,000,000.00 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement | Achievement of Specified Research, Clinical Development, Regulatory and First Commercial Sale Milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | 1,260,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement | Achievement of Specified Sales-based Milestones if Annual Worldwide Net Sales of Licensed Products Reach Specified Levels |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received | $ 1,750,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  | $ 12,000,000.0 |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  | $ 12,000,000.0 |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  | 25,000,000.0 |  |  |  | $ 70,000,000.0 |  | 1,000,000.0 |  |  | 25,000,000.0 |
Deferred revenue |  |  |  | 1,000,000.0 |  | 4,000,000.0 |  |  |  |  |  | 1,000,000.0 |  |  | 1,000,000.0 |
Contract to perform for others, cumulative compensation earned |  |  |  | 24,700,000 |  |  |  |  |  |  |  | 24,700,000 |  |  | 24,700,000 |
Revenue reduction due to decrease in measure of proportional cumulative performance |  |  |  |  |  |  |  |  |  |  |  |  | 3,000,000.0 |  |  |
Increase (decrease) in revenue |  |  |  | 2,400,000 |  |  |  |  |  |  |  |  |  |  |  |
Increase (decrease) in net loss |  |  |  | $ 2,400,000 |  |  |  |  |  |  |  |  |  |  |  |
Increase (decrease) in basic net loss per share (in dollars per share) | $ / shares |  |  |  | $ 0.02 |  |  |  |  |  |  |  |  |  |  |  |
Pfizer | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | 360,000 | 615,000 |  |  |
Pfizer | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | $ 7,600,000 |  | 8,000,000.0 |  |  |  |  |  | 7,600,000 |  |  | 7,600,000 |
Number of milestones included in transaction price | milestone |  |  |  |  |  |  |  |  | 0 |  |  |  |  |  |  |
Pfizer | Maximum | SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  |  |  |  |  |  | 266,500,000 |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  | 300,000,000.0 |  |  |  |  |  |
Pfizer | Maximum | Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  | 175,000,000.0 |  |  |  |  |  |
Pfizer | Achievement of Specified Clinical Development Intellectual Property and Regulatory Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  | 208,500,000 |  |  |  |  |  |
Pfizer | Achievement of First Commercial Sale Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  | $ 475,000,000.0 |  |  |  |  |  |
Pfizer | Achievement of Specified Preclinical Development Clinical Development and First Commercial Sale Milestones | Maximum | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  | $ 60,000,000.0 |  |  |  |  |  |  |
Pfizer | Achievement of Commercial Milestones | Maximum | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  | $ 90,000,000.0 |  |  |  |  |  |  |
Pfizer SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  | $ 104,000,000.0 |  |  |  |
Number of products approved | product |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |
Research Service Fees |  |  |  |  |  |  |  |  |  |  |  | $ 79,000,000.0 |  |  |  |
Agreement termination, term |  |  |  |  |  |  |  |  |  |  |  | 15 years |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | $ 3,183,000 | (3,040,000) |  |  |
Pfizer SB-525 | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | $ 2,190,000 | (3,040,000) |  |  |
Pfizer SB-525 | SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of milestones included in transaction price | milestone |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |
Pfizer SB-525 | Amended Collaboration And License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 300,000 |  |  |  |  |  |  |  | $ 300,000 |  |  | 300,000 |
Milestone revenue receivable |  |  |  | 25,000,000.0 |  |  |  |  |  |  |  | $ 25,000,000.0 |  |  | 25,000,000.0 |
Sanofi |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  | $ 20,000,000.0 |  |  |  |  |
Number of products approved | product |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |
Agreement termination, term |  |  |  |  |  |  |  |  |  |  |  | 180 days |  |  |  |
Increase (decrease) in revenue |  |  |  | (2,200,000) |  |  |  |  |  |  |  |  |  |  |  |
Increase (decrease) in net loss |  |  |  | $ (2,200,000) |  |  |  |  |  |  |  |  |  |  |  |
Increase (decrease) in basic net loss per share (in dollars per share) | $ / shares |  |  |  | $ (0.02) |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | $ 499,000 | 1,949,000 |  |  |
Number of research programs | program |  |  |  |  |  |  |  |  |  |  | 2 |  |  |  |  |
Sanofi | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  | (729,000) | $ 752,000 |  |  |
Sanofi | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  | $ 93,300,000 |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  | 20,000,000.0 |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs |  |  |  |  |  |  |  |  |  |  | 59,800,000 |  |  |  |  |
Deferred revenue |  |  |  | $ 2,900,000 |  | 1,700,000 |  |  |  |  |  | $ 2,900,000 |  |  | 2,900,000 |
Number of milestones included in transaction price | milestone |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |
Cumulative milestone achieved |  |  |  | 6,800,000 |  |  |  |  |  |  |  | $ 6,800,000 |  |  | 6,800,000 |
Sanofi | Collaboration and License Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  | 276,300,000 |  |  |  |  |
Sanofi | Collaboration and License Agreement | Milestone Two |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  | $ 7,500,000 |  |  |  |  |  |  |  |  |  |
Milestone revenue reversal |  |  |  | 300,000 |  |  |  |  |  |  |  | 300,000 |  |  | 300,000 |
Sanofi | Collaboration and License Agreement | Milestone Three |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  | 13,500,000 |  |  |  |  |  |  |  | 13,500,000 |  |  | 13,500,000 |
Sanofi | Collaboration and License Agreement | ST-40 Beta Thalassemia Phase 1 Clinical Trial Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  | 6,000,000.0 |  |  |  |
Cumulative milestone achieved |  |  |  | 5,500,000 |  |  |  |  |  |  |  | 5,500,000 |  |  | 5,500,000 |
Milestone revenue reversal |  |  |  | $ 200,000 |  |  |  |  |  |  |  | $ 200,000 |  |  | $ 200,000 |
Sanofi | Collaboration and License Agreement | Achievement Of Specified Clinical Development And Regulatory Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  | 115,800,000 |  |  |  |  |
Sanofi | Collaboration and License Agreement | Achievement of Specified Sales Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  | $ 160,500,000 |  |  |  |  |

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