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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
$ in Thousands
Total
Revision of Prior Period, Accounting Standards Update, Adjustment
Common Stock
Additional Paid-in Capital
Accumulated Deficit
Accumulated Deficit
Revision of Prior Period, Accounting Standards Update, Adjustment
Accumulated Other Comprehensive Loss
Non- Controlling Interest
Beginning Balances (in shares) at Dec. 31, 2018     102,187,471          
Beginning Balances at Dec. 31, 2018 $ 367,257 $ 897 $ 1,022 $ 929,632 $ (562,696) $ 897 $ (1,440) $ 739
Increase (Decrease) in Stockholders' Equity [Roll Forward]                
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)     633,916          
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 2,660   $ 6 2,654        
Issuance of common stock under employee stock purchase plan (in shares)     131,709          
Issuance of common stock under employee stock purchase plan 1,138   $ 1 1,137        
Issuance of common stock under public offering, net of issuance costs (in shares)     12,650,000          
Issuance of common stock under public offering, net of issuance costs 136,566   $ 127 136,439        
Issuance costs related to public offering (276)     (276)        
Stock-based compensation 9,390     9,390        
Foreign currency translation adjustment (62)              
Net unrealized gain on marketable securities 737           737  
Net loss (72,559)       (72,434)     (125)
Ending Balances (in shares) at Jun. 30, 2019     115,603,096          
Ending Balances at Jun. 30, 2019 445,748   $ 1,156 1,078,976 (634,233)   (765) 614
Beginning Balances (in shares) at Mar. 31, 2019     102,328,752          
Beginning Balances at Mar. 31, 2019 329,181   $ 1,023 934,112 (603,949)   (2,691) 686
Increase (Decrease) in Stockholders' Equity [Roll Forward]                
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)     492,635          
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 2,444   $ 5 2,439        
Issuance of common stock under employee stock purchase plan (in shares)     131,709          
Issuance of common stock under employee stock purchase plan $ 1,138   $ 1 1,137        
Issuance of common stock under public offering, net of issuance costs (in shares) 12,650,000              
Issuance of common stock under public offering, net of issuance costs $ 136,566   $ 127 136,439        
Issuance costs related to public offering (18)     (18)        
Stock-based compensation 4,867     4,867        
Foreign currency translation adjustment 1,442           1,442  
Net unrealized gain on marketable securities 484           484  
Net loss (30,356)       (30,284)     (72)
Ending Balances (in shares) at Jun. 30, 2019     115,603,096          
Ending Balances at Jun. 30, 2019 445,748   $ 1,156 1,078,976 (634,233)   (765) 614
Beginning Balances (in shares) at Dec. 31, 2019     115,972,708          
Beginning Balances at Dec. 31, 2019 432,739   $ 1,160 1,090,828 (656,985)   (2,449) 185
Increase (Decrease) in Stockholders' Equity [Roll Forward]                
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)     409,107          
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 852   $ 4 848        
Issuance of common stock under employee stock purchase plan (in shares)     171,305          
Issuance of common stock under employee stock purchase plan 1,187   $ 2 1,185        
Issuance of common stock under public offering, net of issuance costs (in shares)     24,420,157          
Issuance of common stock under public offering, net of issuance costs 142,526   $ 244 142,282        
Acquisition of additional shares of Sangamo France (339)              
Stock-based compensation 12,384     12,384        
Foreign currency translation adjustment 150           150  
Net unrealized gain on marketable securities 646           646  
Net loss (78,939)       (78,842)     (97)
Ending Balances (in shares) at Jun. 30, 2020     140,973,277          
Ending Balances at Jun. 30, 2020 511,206   $ 1,410 1,247,527 (735,827)   (1,653) (251)
Beginning Balances (in shares) at Mar. 31, 2020     116,278,553          
Beginning Balances at Mar. 31, 2020 394,415   $ 1,163 1,096,854 (699,898)   (3,828) 124
Increase (Decrease) in Stockholders' Equity [Roll Forward]                
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)     103,262          
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 443   $ 1 442        
Issuance of common stock under employee stock purchase plan (in shares)     171,305          
Issuance of common stock under employee stock purchase plan 1,187   $ 2 1,185        
Issuance of common stock under public offering, net of issuance costs (in shares)     24,420,157          
Issuance of common stock under public offering, net of issuance costs 142,526   $ 244 142,282        
Acquisition of additional shares of Sangamo France (339)             (339)
Stock-based compensation 6,764     6,764        
Foreign currency translation adjustment 1,783           1,783  
Net unrealized gain on marketable securities 392           392  
Net loss (35,965)       (35,929)     (36)
Ending Balances (in shares) at Jun. 30, 2020     140,973,277          
Ending Balances at Jun. 30, 2020 $ 511,206   $ 1,410 $ 1,247,527 $ (735,827)   $ (1,653) $ (251)