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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($) $ in Thousands | Total | Cumulative Effect, Period of Adoption, Adjustment | Common Stock | Additional Paid-in Capital | Accumulated Deficit | Accumulated Deficit Cumulative Effect, Period of Adoption, Adjustment | Accumulated Other Comprehensive Income | Non- Controlling Interest
--- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning Balances (in shares) at Dec. 31, 2018 |  |  | 102,187,471 |  |  |  |  |
Beginning Balances at Dec. 31, 2018 | $ 367,257 | $ 897 | $ 1,022 | $ 929,632 | $ (562,696) | $ 897 | $ (1,440) | $ 739
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  |  | 800,818 |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 3,363 |  | $ 8 | 3,355 |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  |  | 131,709 |  |  |  |  |
Issuance of common stock under employee stock purchase plan | 1,139 |  | $ 1 | 1,138 |  |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  |  | 12,650,000 |  |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 136,308 |  | $ 127 | 136,181 |  |  |  |
Acquisition of additional shares of Sangamo France | (321) |  |  |  |  |  |  | (321)
Issuance costs related to Sangamo France Acquisition | (25) |  |  | (25) |  |  |  |
Stock-based compensation | 14,091 |  |  | 14,091 |  |  |  |
Foreign currency translation adjustment | (4,139) |  |  |  |  |  | (4,139) |
Net unrealized loss on marketable securities | 678 |  |  |  |  |  | 678 |
Net loss | (99,920) |  |  |  | (99,741) |  |  | (179)
Ending Balances (in shares) at Sep. 30, 2019 |  |  | 115,769,998 |  |  |  |  |
Ending Balances at Sep. 30, 2019 | 419,328 |  | $ 1,158 | 1,084,372 | (661,540) |  | (4,901) | 239
Beginning Balances (in shares) at Jun. 30, 2019 |  |  | 115,603,096 |  |  |  |  |
Beginning Balances at Jun. 30, 2019 | 445,748 |  | $ 1,156 | 1,078,976 | (634,233) |  | (765) | 614
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  |  | 166,902 |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 704 |  | $ 2 | 702 |  |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  |  |  |  |  |  |  |
Issuance of common stock under public offering, net of issuance costs | (321) |  |  |  |  |  |  | (321)
Issuance costs related to Sangamo France Acquisition | (7) |  |  | (7) |  |  |  |
Stock-based compensation | 4,701 |  |  | 4,701 |  |  |  |
Foreign currency translation adjustment | (4,077) |  |  |  |  |  | (4,077) |
Net unrealized loss on marketable securities | (59) |  |  |  |  |  | (59) |
Net loss | (27,361) |  |  |  | (27,307) |  |  | (54)
Ending Balances (in shares) at Sep. 30, 2019 |  |  | 115,769,998 |  |  |  |  |
Ending Balances at Sep. 30, 2019 | 419,328 |  | $ 1,158 | 1,084,372 | (661,540) |  | (4,901) | 239
Beginning Balances (in shares) at Dec. 31, 2019 |  |  | 115,972,708 |  |  |  |  |
Beginning Balances at Dec. 31, 2019 | 432,739 |  | $ 1,160 | 1,090,828 | (656,985) |  | (2,449) | 185
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  |  | 648,660 |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 2,468 |  | $ 6 | 2,462 |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) |  |  | 171,305 |  |  |  |  |
Issuance of common stock under employee stock purchase plan | 1,187 |  | $ 2 | 1,185 |  |  |  |
Issuance of common stock under public offering, net of issuance costs (in shares) |  |  | 24,420,157 |  |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 142,526 |  | $ 244 | 142,282 |  |  |  |
Acquisition of additional shares of Sangamo France | (660) |  |  |  |  |  |  | (660)
Stock-based compensation | 19,066 |  |  | 19,066 |  |  |  |
Foreign currency translation adjustment | 3,989 |  |  |  |  |  | 3,989 |
Net unrealized loss on marketable securities | 13 |  |  |  |  |  | 13 |
Net loss | (80,447) |  |  |  | (80,392) |  |  | (55)
Ending Balances (in shares) at Sep. 30, 2020 |  |  | 141,212,830 |  |  |  |  |
Ending Balances at Sep. 30, 2020 | 520,881 |  | $ 1,412 | 1,255,823 | (737,377) |  | 1,553 | (530)
Beginning Balances (in shares) at Jun. 30, 2020 |  |  | 140,973,277 |  |  |  |  |
Beginning Balances at Jun. 30, 2020 | 511,206 |  | $ 1,410 | 1,247,527 | (735,827) |  | (1,653) | (251)
Increase (Decrease) in Stockholders' Equity [Roll Forward] |  |  |  |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  |  | 239,553 |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | 1,616 |  | $ 2 | 1,614 |  |  |  |
Acquisition of additional shares of Sangamo France | (321) |  |  |  |  |  |  | (321)
Stock-based compensation | 6,682 |  |  | 6,682 |  |  |  |
Foreign currency translation adjustment | 3,839 |  |  |  |  |  | 3,839 |
Net unrealized loss on marketable securities | (633) |  |  |  |  |  | (633) |
Net loss | (1,508) |  |  |  | (1,550) |  |  | 42
Ending Balances (in shares) at Sep. 30, 2020 |  |  | 141,212,830 |  |  |  |  |
Ending Balances at Sep. 30, 2020 | $ 520,881 |  | $ 1,412 | $ 1,255,823 | $ (737,377) |  | $ 1,553 | $ (530)

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