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MAJOR CUSTOMERS, PARTNERSHIPS AND STRATEGIC ALLIANCES - Additional Information (Detail) |  |  | 1 Months Ended | 3 Months Ended | 9 Months Ended | 41 Months Ended | 81 Months Ended |
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Jul. 27, 2020 USD ($) | Apr. 05, 2018 USD ($) | Oct. 30, 2020 USD ($) | Sep. 30, 2020 USD ($) milestone | Aug. 31, 2020 USD ($) | May 31, 2020 USD ($) | Apr. 30, 2020 USD ($) $ / shares shares | Mar. 31, 2020 USD ($) $ / shares | Feb. 29, 2020 USD ($) product_target | Dec. 31, 2019 USD ($) | Sep. 30, 2019 USD ($) | Aug. 31, 2019 USD ($) | May 31, 2018 USD ($) | Dec. 31, 2017 USD ($) | May 31, 2017 USD ($) | Jan. 31, 2014 USD ($) program | Sep. 30, 2020 USD ($) milestone product_target | Sep. 30, 2019 USD ($) | Sep. 30, 2020 USD ($) milestone option product $ / shares | Sep. 30, 2019 USD ($) | Sep. 30, 2020 USD ($) milestone | Sep. 30, 2020 USD ($) product milestone | Feb. 28, 2020
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 57,763,000 | $ 21,958,000 | $ 92,392,000 | $ 47,577,000 |  |  |
Maximum number of product targets replaced | product_target |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 10 |  |  |  |  |  |
Recognized portion of equity issued |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,000 |  | $ 25,000 |  |  |
Revenue reduction due to decrease in measure of proportional cumulative performance, percentage |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 38.00% |  |  |
Other long-term liabilities |  |  |  | $ 6,238,000 |  |  |  |  |  | $ 5,711,000 |  |  |  |  |  |  | $ 6,238,000 |  | 6,238,000 |  | $ 6,238,000 | $ 6,238,000 |
California Institute For Regenerative Medicine Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Funds due under the agreement |  |  |  |  |  |  |  |  |  |  |  |  | $ 8,000,000.0 |  |  |  |  |  |  |  |  |  |
Other long-term liabilities |  |  |  | 6,200,000 |  |  |  |  |  | 5,700,000 |  |  |  |  |  |  | 6,200,000 |  | 6,200,000 |  | 6,200,000 | 6,200,000 |
California Institute For Regenerative Medicine Agreement | Research Grants |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5,200,000 |  |  |  |
Novartis | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Proceeds from collaborators |  |  |  |  | $ 75,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price | $ 95,100,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
License fee | 75,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs | 20,100,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 75,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  | 75,000,000.0 |  | 75,000,000.0 |  | 75,000,000.0 | 75,000,000.0 |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  | 0 |  |  |  |
Collaboration Arrangement, Commission Fee, Portion Of Gross Proceeds, Value |  |  |  | 1,500,000 |  |  |  |  |  |  |  |  |  |  |  |  | $ 1,500,000 |  | 1,500,000 |  | 1,500,000 | 1,500,000 |
Percent of initial recognition |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2.00% |  |  |  |  |  |
Novartis | Collaboration and License Agreement | Achievement of Specified Preclinical Development Clinical Development and First Commercial Sale Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received | 420,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Novartis | Collaboration and License Agreement | Achievement of Commercial Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received | $ 300,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 9,621,000 | 0 | 17,799,000 | $ 0 |  |  |
Biogen MA, Inc. | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,306,000 | 0 | 14,050,000 | 0 |  |  |
Biogen MA, Inc. | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Proceeds from collaborators |  |  |  |  |  | $ 125,000,000.0 |  |  | $ 125,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  |  |  |  |  | $ 2,370,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of product targets | product_target |  |  |  |  |  |  |  |  | 12 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of product targets selected | product_target |  |  |  |  |  |  |  |  | 3 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of additional product targets | product_target |  |  |  |  |  |  |  |  | 9 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Target selection period |  |  |  |  |  |  |  |  | 5 years |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaboration Arrangement, Commission Fee, Portion Of Gross Proceeds, Value |  |  |  | 7,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  | $ 7,000,000.0 |  | $ 7,000,000.0 |  | 7,000,000.0 | 7,000,000.0 |
Percent of initial recognition |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2.00% |  | 2.00% |  |  |  |
Portion of contract asset recognized |  |  |  | 4,100,000 |  |  |  |  |  |  |  |  |  |  |  |  | $ 4,100,000 |  | $ 4,100,000 |  | 4,100,000 | 4,100,000 |
Amortization |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 100,000 |  | 300,000 |  |  |  |
Biogen MA, Inc. | Collaboration and License Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research period |  |  |  |  |  |  |  |  | 7 years |  |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Collaboration and License Agreement | Pre-approval Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  |  |  |  |  | $ 925,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Collaboration and License Agreement | Sales-based Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  |  |  |  |  | $ 1,450,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Stock Purchase Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 190,500,000 |  |  |  |  |  |  |  |  |  |  |  |  | 190,500,000 |  | 190,500,000 |  | 190,500,000 | 190,500,000 |
Number of shares issued in transaction (in shares) | shares |  |  |  |  |  |  | 24,420,157 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Sale of stock, price per share (in dollars per share) | $ / shares |  |  |  |  |  |  | $ 9.2137 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Consideration received on transaction |  |  |  |  |  |  | $ 225,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Standstill restriction period |  |  |  |  |  |  |  |  | 3 years |  |  |  |  |  |  |  |  |  |  |  |  |  |
Standstill restriction, ownership threshold percentage ownership percentage |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5.00%
Agreement restriction, percentage of shares held |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 50.00%
Voting provisions expiration period |  |  |  |  |  |  |  |  | 2 years |  |  |  |  |  |  |  |  |  |  |  |  |  |
Voting provisions, ownership threshold percentage |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5.00%
Excess consideration received on transaction |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 79,600,000 |  |  |  |
Recognized portion of equity issued |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2,900,000 |  |  |  |  |  |
Collaboration agreement, equity issued |  |  |  | 145,400,000 |  |  |  |  |  |  |  |  |  |  |  |  | 145,400,000 |  | 145,400,000 |  | 145,400,000 | 145,400,000 |
Kite Pharma Inc |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,294,000 | 8,861,000 | 21,935,000 | 26,233,000 |  |  |
Kite Pharma Inc | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 6,296,000 | 6,296,000 | 18,750,000 | 18,682,000 |  |  |
Kite Pharma Inc | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  | $ 189,300,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs |  | 39,300,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 87,700,000 |  |  |  |  |  | 106,500,000 |  |  |  |  |  |  | 87,700,000 |  | $ 87,700,000 |  | 87,700,000 | 87,700,000 |
Milestone payments received |  | $ 150,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Initial research term of agreement |  | 6 years |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of options to extend initial research term | option |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2 |  |  |  |
Extended research term of agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 1 year |  |  |  |
Separate upfront fee |  | $ 10,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs for new research plan |  |  |  |  |  |  |  |  |  |  | $ 3,400,000 |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  | 150,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  | 3,010,000,000.00 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement | Achievement of Specified Research, Clinical Development, Regulatory and First Commercial Sale Milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received |  | 1,260,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Kite Pharma Inc | Collaboration and License Agreement | Achievement of Specified Sales-based Milestones if Annual Worldwide Net Sales of Licensed Products Reach Specified Levels |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received |  | $ 1,750,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 700,000 |  |  |  |  |  | 4,000,000.0 |  |  |  |  |  |  | 700,000 |  | $ 700,000 |  | 700,000 | 700,000 |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 9,200,000 | 468,000 | 12,985,000 | 1,538,000 |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 12,000,000.0 |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  | 25,000,000.0 |  |  |  |  |  |  |  |  | 1,200,000 |  |  |  |
Contract to perform for others, cumulative compensation earned |  |  |  | 24,800,000 |  |  |  |  |  |  |  |  |  |  |  |  | 24,800,000 |  | 24,800,000 |  | 24,800,000 | 24,800,000 |
Revenue reduction due to decrease in measure of proportional cumulative performance |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 3,000,000.0 |  |  |
Increase (decrease) in revenue |  |  |  |  |  |  |  | $ 2,400,000 |  |  |  |  |  |  |  |  |  |  | 8,800,000 |  |  |  |
Increase (decrease) in net loss |  |  |  |  |  |  |  | $ 2,400,000 |  |  |  |  |  |  |  |  |  |  | $ 8,800,000 |  |  |  |
Increase (decrease) in basic net loss per share (in dollars per share) | $ / shares |  |  |  |  |  |  |  | $ (0.02) |  |  |  |  |  |  |  |  |  |  | $ 0.07 |  |  |  |
Pfizer | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 4,200,000 | 468,000 | $ 7,985,000 | 1,538,000 |  |  |
Pfizer | SB-525 and Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 70,000,000.0 |  |  |  |  |  |  |  |
Pfizer | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  | 5,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  |  |  | 17,000,000.0 |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | $ 0.0 |  |  |  |  |  | 8,000,000.0 |  |  |  |  |  |  | 0.0 |  | 0.0 |  | 0.0 | 0.0 |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  |  |  | 12,000,000.0 |  |  |  |  |  |  |  |  |
Extended research term of agreement |  |  |  | 2 years |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  | 5,000,000.0 |  |  | 5,000,000.0 |  | 5,000,000.0 |  |  |  |
Pfizer | Milestone Achievement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5,000,000 | 0 | 5,000,000 | 0 |  |  |
Pfizer | Maximum | SB-525 and Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 266,500,000 |  |  |  |  |  |  |  |
Pfizer | Maximum | SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 300,000,000.0 |  |  |  |  |  |  |  |
Pfizer | Maximum | Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 175,000,000.0 |  |  |  |  |  |  |  |
Pfizer | Phase 3 Clinical Trial |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contract to perform for others, cumulative compensation earned |  |  |  | $ 29,800,000 |  |  |  |  |  |  |  |  |  |  |  |  | 29,800,000 |  | 29,800,000 |  | 29,800,000 | $ 29,800,000 |
Pfizer | Phase 3 Clinical Trial | Forecast | Subsequent Event |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  | $ 30,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Pfizer | Achievement of Specified Clinical Development Intellectual Property and Regulatory Milestones | Maximum | SB-525 and Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 208,500,000 |  |  |  |  |  |  |  |
Pfizer | Achievement of First Commercial Sale Milestones | Maximum | SB-525 and Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 475,000,000.0 |  |  |  |  |  |  |  |
Pfizer | Achievement of Specified Preclinical Development Clinical Development and First Commercial Sale Milestones | Maximum | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  | 60,000,000.0 |  |  |  |  |  |  |  |  |
Pfizer | Achievement of Commercial Milestones | Maximum | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 90,000,000.0 |  |  |  |  |  |  |  |  |
Pfizer SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 134,000,000.0 |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 30,084,000 | 3,440,000 | $ 33,696,000 | 5,035,000 |  |  |
Agreement termination, term |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 15 years |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 55,000,000.0 |  |  |  |
Research Service Fees |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 79,000,000.0 |  |  |  |
Pfizer SB-525 | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 221,000 | 3,440,000 | $ 2,737,000 | 5,035,000 |  |  |
Pfizer SB-525 | SB-525 and Other Products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 55,000,000.0 |  |
Number of milestones achieved | milestone |  |  |  | 2 |  |  |  |  |  |  |  |  |  |  |  |  | 2 |  | 2 |  | 2 | 2 |
Number of products approved | product |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |
Number of milestones included in transaction price | milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |
Pfizer SB-525 | Milestone Achievement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 29,863,000 | 0 | $ 30,959,000 | 0 |  |  |
Pfizer SB-525 | Amended Collaboration And License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | $ 200,000 |  |  |  |  |  |  |  |  |  |  |  |  | 200,000 |  | 200,000 |  | $ 200,000 | $ 200,000 |
Pfizer SB-525 | Amended Collaboration And License Agreement | Phase 3 Clinical Trial |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  | 200,000 |  |  |  |  |  |  |  |  |  |  |  |  | 200,000 |  | 200,000 |  | 200,000 | $ 200,000 |
Sanofi |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 1,498,000 | 2,754,000 | $ 3,789,000 | 7,763,000 |  |  |
Agreement termination, term |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 180 days |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 20,000,000.0 |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  | 6,000,000.0 |  |  |  |  |  |  | 6,000,000.0 |  | 6,000,000.0 |  |  |
Number of products approved | product |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |
Increase (decrease) in revenue |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ (2,200,000) |  |  |  |
Increase (decrease) in net loss |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 2,200,000 |  |  |  |
Increase (decrease) in basic net loss per share (in dollars per share) | $ / shares |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ (0.02) |  |  |  |
Number of research programs | program |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2 |  |  |  |  |  |  |
Sanofi | Collaboration Agreements |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 321,000 | 940,000 | $ (28,000) | 2,594,000 |  |  |
Sanofi | Milestone Achievement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 216,000 | 0 | (19,000) | 0 |  |  |
Sanofi | Collaboration and License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 93,300,000 |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 59,800,000 |  |  |  |  |  |  |
Deferred revenue |  |  |  | 1,700,000 |  |  |  |  |  | $ 1,700,000 |  |  |  |  |  |  | 1,700,000 |  | 1,700,000 |  | 1,700,000 | $ 1,700,000 |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 20,000,000.0 |  |  |  |  |  |  |
Number of milestones included in transaction price | milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |
Cumulative milestone achieved |  |  |  | 7,100,000 |  |  |  |  |  |  |  |  |  |  |  |  | 7,100,000 |  | 7,100,000 |  | 7,100,000 | $ 7,100,000 |
Sanofi | Collaboration and License Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 276,300,000 |  |  |  |  |  |  |
Sanofi | Collaboration and License Agreement | Milestone Two |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  | $ 7,500,000 |  |  |  |  |  |  | $ 7,500,000 |  | $ 7,500,000 |  |  |
Sanofi | Collaboration and License Agreement | Milestone Three |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  | 13,500,000 |  |  |  |  |  |  |  |  |  |  |  |  | 13,500,000 |  | 13,500,000 |  | 13,500,000 | 13,500,000 |
Sanofi | Collaboration and License Agreement | ST-40 Beta Thalassemia Phase 1 Clinical Trial Milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  | $ 6,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Cumulative milestone achieved |  |  |  | $ 5,700,000 |  |  |  |  |  |  |  |  |  |  |  |  | $ 5,700,000 |  | $ 5,700,000 |  | $ 5,700,000 | $ 5,700,000 |
Sanofi | Collaboration and License Agreement | Achievement Of Specified Clinical Development And Regulatory Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 115,800,000 |  |  |  |  |  |  |
Sanofi | Collaboration and License Agreement | Achievement of Specified Sales Milestones | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 160,500,000 |  |  |  |  |  |  |

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