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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 9 Months Ended
Sep. 30, 2020
Sep. 30, 2019
Operating Activities:    
Net loss $ (80,447) $ (99,920)
Adjustments to reconcile net loss to net cash provided by (used in) operating activities:    
Depreciation and amortization 3,999 2,742
Amortization of discount on marketable securities (1,344) (3,715)
Amortization and other changes in operating lease right-of-use assets 5,706 3,796
Gain on free shares (31) (488)
Stock-based compensation 19,066 14,091
Loss on disposal of property and equipment 197 0
Net loss on lease termination 0 218
Other 0 (29)
Net changes in operating assets and liabilities:    
Interest receivable (140) (305)
Accounts receivable (2,578) (17,280)
Prepaid expenses and other assets (9,098) (4,734)
Accounts payable and accrued liabilities 1,336 (1,753)
Accrued compensation and employee benefits 4,186 1,608
Deferred revenues 235,551 (20,477)
Long-term portion of lease liabilities (2,760) (920)
Other non-current liabilities 527 3,580
Net cash provided by (used in) operating activities 174,170 (123,586)
Investing Activities:    
Purchases of marketable securities (335,002) (321,390)
Maturities of marketable securities 205,039 292,147
Purchases of property and equipment (10,703) (13,894)
Purchase of additional shares of Sangamo France (503) (262)
Net cash used in investing activities (141,169) (43,399)
Financing Activities:    
Proceeds from public offering of common stock, net of issuance costs 0 136,308
Proceeds from issuance of common stock in connection with the Biogen collaboration agreement, net of issuance costs 142,526 0
Taxes paid related to net share settlement of equity awards (573) (388)
Proceeds from exercise of stock options and restricted stock units 3,041 3,751
Proceeds from exercise of stock options and restricted stock units 1,187 1,139
Net cash provided by financing activities 146,181 140,810
Effect of exchange rate changes on cash, cash equivalents, and restricted cash (217) 48
Net increase (decrease) in cash, cash equivalents, and restricted cash 178,965 (26,127)
Cash, cash equivalents, and restricted cash, beginning of period 81,928 143,918
Cash, cash equivalents, and restricted cash, end of period 260,893 117,791
Supplemental disclosure of non-cash activities:    
Property and equipment included in unpaid liabilities 899 4,257
Right-of-use assets obtained in exchange for lease obligations $ 0 $ 29,647