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Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Total | Cumulative Effect, Period of Adoption, Adjustment | Common Stock | Additional Paid-in Capital | Accumulated Deficit | Accumulated Deficit Cumulative Effect, Period of Adoption, Adjustment | Accumulated Other Comprehensive Income (Loss) | Non- Controlling Interest
--- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning Balances (in shares) at Dec. 31, 2017 |  |  | 85,598,534 |  |  |  |  |
Beginning Balances at Dec. 31, 2017 | $ 187,900 | $ 1,117 | $ 856 | $ 682,809 | $ (495,479) | $ 1,117 | $ (286) | $ 0
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  |  | 2,103,727 |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | $ 14,467 |  | $ 20 | 14,447 |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) | 328,710 |  | 328,710 |  |  |  |  |
Issuance of common stock under employee stock purchase plan | $ 1,484 |  | $ 4 | 1,480 |  |  |  |
Issuance of common stock, net of issuance costs (in shares) |  |  | 14,156,500 |  |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 215,758 |  | $ 142 | 215,616 |  |  |  |
Stock-based compensation | 14,677 |  |  | 14,677 |  |  |  |
Acquisition of additional shares of Sangamo France | 603 |  |  | 603 |  |  |  |
Non-controlling interest upon acquisition of Sangamo France | 1,313 |  |  |  |  |  |  | 1,313
Foreign currency translation adjustment | (1,148) |  |  |  |  |  | (1,129) | (19)
Net pension losses | (21) |  |  |  |  |  | (21) |
Net unrealized loss on marketable securities, net of tax | (4) |  |  |  |  |  | (4) |
Net loss | (68,889) |  |  |  | (68,334) |  |  | (555)
Ending Balances (in shares) at Dec. 31, 2018 |  |  | 102,187,471 |  |  |  |  |
Ending Balances at Dec. 31, 2018 | $ 367,257 | $ 897 | $ 1,022 | 929,632 | (562,696) | $ 897 | (1,440) | 739
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Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  |  | 885,873 |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | $ 3,677 |  | $ 9 | 3,668 |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) | 249,364 |  | 249,364 |  |  |  |  |
Issuance of common stock under employee stock purchase plan | $ 2,044 |  | $ 2 | 2,042 |  |  |  |
Issuance of common stock, net of issuance costs (in shares) |  |  | 12,650,000 |  |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 136,308 |  | $ 127 | 136,181 |  |  |  |
Stock-based compensation | 19,330 |  |  | 19,330 |  |  |  |
Acquisition of additional shares of Sangamo France | (321) |  |  |  |  |  |  | (321)
Issuance costs related to Sangamo France acquisition | (25) |  |  | (25) |  |  |  |
Foreign currency translation adjustment | (1,573) |  |  |  |  |  | (1,573) |
Net pension losses | (28) |  |  |  |  |  | (28) |
Net unrealized loss on marketable securities, net of tax | 592 |  |  |  |  |  | 592 |
Net loss | $ (95,419) |  |  |  | (95,186) |  |  | (233)
Ending Balances (in shares) at Dec. 31, 2019 | 115,972,708 |  | 115,972,708 |  |  |  |  |
Ending Balances at Dec. 31, 2019 | $ 432,739 |  | $ 1,160 | 1,090,828 | (656,985) |  | (2,449) | 185
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Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares) |  |  | 1,395,956 |  |  |  |  |
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax | $ 8,559 |  | $ 14 | 8,545 |  |  |  |
Issuance of common stock under employee stock purchase plan (in shares) | 274,382 |  | 274,382 |  |  |  |  |
Issuance of common stock under employee stock purchase plan | $ 2,015 |  | $ 3 | 2,012 |  |  |  |
Issuance of common stock, net of issuance costs (in shares) |  |  | 24,420,157 |  |  |  |  |
Issuance of common stock under public offering, net of issuance costs | 142,526 |  | $ 244 | 142,282 |  |  |  |
Stock-based compensation | 25,708 |  |  | 25,708 |  |  |  |
Acquisition of additional shares of Sangamo France | (927) |  |  |  |  |  |  | (927)
Foreign currency translation adjustment | 8,345 |  |  |  |  |  | (8,345) |
Net pension losses | (193) |  |  |  |  |  | (193) |
Net unrealized loss on marketable securities, net of tax | (284) |  |  |  |  |  | (284) |
Net loss | $ (121,122) |  |  |  | (120,996) |  |  | (126)
Ending Balances (in shares) at Dec. 31, 2020 | 142,063,203 |  | 142,063,203 |  |  |  |  |
Ending Balances at Dec. 31, 2020 | $ 497,366 |  | $ 1,421 | $ 1,269,375 | $ (777,981) |  | $ 5,419 | $ (868)

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