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Consolidated Statements of Stockholders' Equity - USD ($)
$ in Thousands
Total
Cumulative Effect, Period of Adoption, Adjustment
Common Stock
Additional Paid-in Capital
Accumulated Deficit
Accumulated Deficit
Cumulative Effect, Period of Adoption, Adjustment
Accumulated Other Comprehensive Income (Loss)
Non- Controlling Interest
Beginning Balances (in shares) at Dec. 31, 2017     85,598,534          
Beginning Balances at Dec. 31, 2017 $ 187,900 $ 1,117 $ 856 $ 682,809 $ (495,479) $ 1,117 $ (286) $ 0
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)     2,103,727          
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax $ 14,467   $ 20 14,447        
Issuance of common stock under employee stock purchase plan (in shares) 328,710   328,710          
Issuance of common stock under employee stock purchase plan $ 1,484   $ 4 1,480        
Issuance of common stock, net of issuance costs (in shares)     14,156,500          
Issuance of common stock under public offering, net of issuance costs 215,758   $ 142 215,616        
Stock-based compensation 14,677     14,677        
Acquisition of additional shares of Sangamo France 603     603        
Non-controlling interest upon acquisition of Sangamo France 1,313             1,313
Foreign currency translation adjustment (1,148)           (1,129) (19)
Net pension losses (21)           (21)  
Net unrealized loss on marketable securities, net of tax (4)           (4)  
Net loss (68,889)       (68,334)     (555)
Ending Balances (in shares) at Dec. 31, 2018     102,187,471          
Ending Balances at Dec. 31, 2018 $ 367,257 $ 897 $ 1,022 929,632 (562,696) $ 897 (1,440) 739
Accounting Standards Update [Extensible List] us-gaap:AccountingStandardsUpdate201409Member              
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)     885,873          
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax $ 3,677   $ 9 3,668        
Issuance of common stock under employee stock purchase plan (in shares) 249,364   249,364          
Issuance of common stock under employee stock purchase plan $ 2,044   $ 2 2,042        
Issuance of common stock, net of issuance costs (in shares)     12,650,000          
Issuance of common stock under public offering, net of issuance costs 136,308   $ 127 136,181        
Stock-based compensation 19,330     19,330        
Acquisition of additional shares of Sangamo France (321)             (321)
Issuance costs related to Sangamo France acquisition (25)     (25)        
Foreign currency translation adjustment (1,573)           (1,573)
Net pension losses (28)           (28)  
Net unrealized loss on marketable securities, net of tax 592           592  
Net loss $ (95,419)       (95,186)     (233)
Ending Balances (in shares) at Dec. 31, 2019 115,972,708   115,972,708          
Ending Balances at Dec. 31, 2019 $ 432,739   $ 1,160 1,090,828 (656,985)   (2,449) 185
Accounting Standards Update [Extensible List] us-gaap:AccountingStandardsUpdate201801Member              
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)     1,395,956          
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax $ 8,559   $ 14 8,545        
Issuance of common stock under employee stock purchase plan (in shares) 274,382   274,382          
Issuance of common stock under employee stock purchase plan $ 2,015   $ 3 2,012        
Issuance of common stock, net of issuance costs (in shares)     24,420,157          
Issuance of common stock under public offering, net of issuance costs 142,526   $ 244 142,282        
Stock-based compensation 25,708     25,708        
Acquisition of additional shares of Sangamo France (927)             (927)
Foreign currency translation adjustment 8,345           (8,345)  
Net pension losses (193)           (193)  
Net unrealized loss on marketable securities, net of tax (284)           (284)  
Net loss $ (121,122)       (120,996)     (126)
Ending Balances (in shares) at Dec. 31, 2020 142,063,203   142,063,203          
Ending Balances at Dec. 31, 2020 $ 497,366   $ 1,421 $ 1,269,375 $ (777,981)   $ 5,419 $ (868)