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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2021
Mar. 31, 2020
Operating Activities:    
Net loss $ (45,939) $ (42,974)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 1,875 1,311
Amortization of premium (discount) on marketable securities 824 (744)
Amortization and other changes in operating lease right-of-use assets 2,023 1,886
Loss on free shares 27 73
Stock-based compensation 7,524 5,620
Net changes in operating assets and liabilities:    
Interest receivable 61 (244)
Accounts receivable (1,522) 29,939
Prepaid expenses and other assets (1,741) (100)
Accounts payable and accrued liabilities (3,829) 345
Accrued compensation and employee benefits (7,561) (4,965)
Deferred revenues (20,047) (8,319)
Long-term portion of lease liabilities (1,045) (887)
Other non-current liabilities 800 169
Net cash used in operating activities (68,550) (18,890)
Investing Activities:    
Purchases of marketable securities (97,935) (43,580)
Maturities of marketable securities 144,369 71,075
Sales of marketable securities 6,870 0
Purchases of property and equipment (7,950) (3,775)
Purchase of additional shares of Sangamo France (65) 0
Net cash provided by investing activities 45,289 23,720
Financing Activities:    
Proceeds from at-the-market offering, net of offering expenses 15,651 0
Taxes paid related to net share settlement of equity awards (2,234) (411)
Proceeds from exercise of stock options and restricted stock units 1,818 820
Net cash provided by financing activities 15,235 409
Effect of exchange rate changes on cash, cash equivalents, and restricted cash (328) 82
Net (decrease) increase in cash, cash equivalents, and restricted cash (8,354) 5,321
Cash, cash equivalents, and restricted cash, beginning of period 132,829 81,928
Cash, cash equivalents, and restricted cash, end of period 124,475 87,249
Supplemental cash flow disclosures:    
Property and equipment included in unpaid liabilities 2,953 1,080
Right-of-use assets obtained in exchange for lease obligations $ 1,356 $ 0