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MAJOR CUSTOMERS, PARTNERSHIPS AND STRATEGIC ALLIANCES - Additional Information (Detail) |  | 1 Months Ended | 3 Months Ended | 6 Months Ended | 12 Months Ended | 40 Months Ended | 50 Months Ended |
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Jul. 27, 2020 USD ($) | Sep. 30, 2020 USD ($) | Aug. 31, 2020 USD ($) | May 31, 2020 USD ($) | Apr. 30, 2020 USD ($) $ / shares shares | Feb. 29, 2020 USD ($) product_target | Sep. 30, 2019 USD ($) | Aug. 31, 2019 USD ($) | May 31, 2018 USD ($) | Apr. 30, 2018 USD ($) milestone | Dec. 31, 2017 USD ($) | May 31, 2017 USD ($) | Jan. 31, 2014 USD ($) program | Jun. 30, 2021 USD ($) | Jun. 30, 2020 USD ($) | Jun. 30, 2021 USD ($) option milestone product_target product royaltyFee | Jun. 30, 2020 USD ($) | Dec. 31, 2020 USD ($) | Mar. 31, 2021 USD ($) product milestone | Jun. 30, 2021 USD ($) milestone | Dec. 31, 2019 USD ($)
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 27,872,000 | $ 21,553,000 | $ 54,152,000 | $ 34,629,000 |  |  |  |
Other long-term liabilities |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,425,000 |  | 7,425,000 |  | $ 7,011,000 |  | $ 7,425,000 |
California institute for regenerative medicine agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Funds due under the agreement |  |  |  |  |  |  |  |  | $ 8,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |
Other long-term liabilities |  |  |  |  |  |  |  |  |  |  |  |  |  | 6,800,000 |  | 6,800,000 |  | 6,400,000 |  | 6,800,000 |
California institute for regenerative medicine agreement | Research grants |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5,200,000 |  |  |  |  |
Novartis Institutes for BioMedical Research, Inc. |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 9,581,000 | 0 | 17,442,000 | 0 |  |  |  |
Novartis Institutes for BioMedical Research, Inc. | Recognition of license and stand-ready fee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,559,000 | 0 | 13,759,000 | 0 |  |  |  |
Novartis Institutes for BioMedical Research, Inc. | Collaboration and license agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Proceeds from collaborators |  |  | $ 75,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 75,000,000.0 |  |  |
Collaborative arrangement transaction price | $ 95,100,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
License fee | 75,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs | 20,100,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  |  |  |  |  |  | 57,100,000 |  | 57,100,000 |  | 70,900,000 |  | 57,100,000 |
Financial advisory fees |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 1,500,000 |  |  |
Percent of initial recognition |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2.00% |  |  |
Amortization |  |  |  |  |  |  |  |  |  |  |  |  |  | 200,000 |  | 300,000 |  |  |  |  |
Novartis Institutes for BioMedical Research, Inc. | Collaboration and license agreement | Achievement of specified preclinical development clinical development and first commercial sale milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received | 420,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Novartis Institutes for BioMedical Research, Inc. | Collaboration and license agreement | Achievement of commercial milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received | $ 300,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 10,767,000 | 8,178,000 | 21,217,000 | 8,178,000 |  |  |  |
Biogen MA, Inc. | Recognition of license and stand-ready fee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,306,000 | 6,744,000 | 14,612,000 | 6,744,000 |  |  |  |
Biogen MA, Inc. | Collaboration and license agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Proceeds from collaborators |  |  |  | $ 125,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 125,000,000.0 |  |  |
Financial advisory fees |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 7,000,000.0 |  |  |
Percent of initial recognition |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2.00% |  |  |
Amortization |  |  |  |  |  |  |  |  |  |  |  |  |  | 200,000 | 0 | $ 300,000 | 0 |  |  |  |
Maximum milestone payment receivable |  |  |  |  |  | $ 2,370,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of product targets | product_target |  |  |  |  |  | 12 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of additional product targets | product_target |  |  |  |  |  | 9 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Target selection period |  |  |  |  |  | 5 years |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of product targets selected | product_target |  |  |  |  |  | 3 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research period |  |  |  |  |  | 7 years |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum number of product targets replaced | product_target |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 10 |  |  |  |  |
Portion of contract asset recognized |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 4,100,000 |  |  |
Consideration received on transaction |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 204,600,000 |  |  |
Biogen MA, Inc. | Collaboration and license agreement | Pre-approval milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  |  | $ 925,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Collaboration and license agreement | Sales-based milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum milestone payment receivable |  |  |  |  |  | $ 1,450,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Biogen MA, Inc. | Stock purchase agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  |  |  |  |  |  | 168,600,000 |  | $ 168,600,000 |  | 183,200,000 |  | 168,600,000 |
Number of shares issued in transaction (in shares) | shares |  |  |  |  | 24,420,157 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Sale of stock, price per share (in dollars per share) | $ / shares |  |  |  |  | $ 9.2137 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Aggregate purchase price |  |  |  |  | $ 204,600,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Standstill restriction period |  |  |  |  |  | 3 years |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Standstill restriction, ownership threshold percentage ownership percentage |  |  |  |  |  | 5.00% |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Agreement restriction, percentage of shares held |  |  |  |  |  | 50.00% |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Voting provisions expiration period |  |  |  |  |  | 2 years |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Voting provisions, ownership threshold percentage |  |  |  |  |  | 5.00% |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Excess consideration received on transaction |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 79,600,000 |  |  |
Consideration received on transaction |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 225,000,000.0 |  |  |
Recognized portion of equity issued |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2,900,000 |  |  |
Collaboration agreement, equity issued |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 145,400,000 |  |  |
Kite Pharma, Inc. |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 6,324,000 | 7,422,000 | 12,612,000 | 14,641,000 |  |  |  |
Kite Pharma, Inc. | Recognition of license and stand-ready fee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 6,227,000 | 6,227,000 | 12,386,000 | 12,454,000 |  |  |  |
Kite Pharma, Inc. | Collaboration and license agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  | $ 3,010,000,000.00 |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  | $ 189,300,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs |  |  |  |  |  |  |  |  |  |  |  |  |  | 39,300,000 |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  |  |  |  |  |  | 69,000,000.0 |  | $ 69,000,000.0 |  | 81,400,000 |  | 69,000,000.0 |
Milestone payments received |  |  |  |  |  |  |  |  |  | 150,000,000.0 |  |  |  |  |  |  |  |  |  |  |
Initial research term of agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 6 years |  |  |  |  |
Number of options to extend initial research term | option |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2 |  |  |  |  |
Extended research term of agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 1 year |  |  |  |  |
Separate upfront fee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 10,000,000.0 |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs for new research plan |  |  |  |  |  |  | $ 3,400,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  | 150,000,000.0 |  |  |  |  |  |  |
Kite Pharma, Inc. | Collaboration and license agreement | Achievement of specified research, clinical development, regulatory and first commercial sale milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  | 1,260,000,000 |  |  |  |  |  |  |  |  |  |  |
Kite Pharma, Inc. | Collaboration and license agreement | Achievement of specified sales-based milestones if annual worldwide net sales of licensed products reach specified levels |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Contingent development - and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  | $ 1,750,000,000 |  |  |  |  |  |  |  |  |  |  |
Maximum amount of achieved milestones to receive payment | milestone |  |  |  |  |  |  |  |  |  | 10 |  |  |  |  |  |  |  |  |  |  |
Pfizer Inc. |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  | $ 12,000,000.0 |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  | $ 0 |  |  |  |  |
Pfizer Inc. | SB-525 and other products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  | $ 70,000,000.0 |  |  |  |  |  |  |  |  |
Potential amount to be funded for achievement of specified commercialized and sales milestones |  |  |  |  |  |  |  |  |  |  |  | 266,500,000 |  |  |  |  |  |  |  |  |
Pfizer Inc. | SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  |  | 300,000,000.0 |  |  |  |  |  |  |  |  |
Pfizer Inc. | Other products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  |  | 175,000,000.0 |  |  |  |  |  |  |  |  |
Pfizer Inc. | Recognition of license and stand-ready fee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 3,400,000 |  | 3,800,000 |  |  |  |
Pfizer Inc. | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 5,000,000.0 |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  | 17,000,000.0 |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  | 12,000,000.0 |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  | $ 5,000,000.0 |  |  |  |  |  |  |  |  | 5,000,000.0 |  |  |  |  |  |  |  |  |  |
Agreement termination, term |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 15 years |  |  |  |  |
Number of products approved | product |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |
Number of milestones included in transaction price | milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |
Pfizer Inc. | Achievement of specified preclinical development clinical development and first commercial sale milestones | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  | 60,000,000.0 |  |  |  |  |  |  |  |  |  |
Pfizer Inc. | Achievement of commercial milestones | C9ORF72 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  | $ 90,000,000.0 |  |  |  |  |  |  |  |  |  |
Pfizer Inc. | Achievement of specified clinical development intellectual property and regulatory milestones | SB-525 and other products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  | 208,500,000 |  |  |  |  |  |  |  |  |
Pfizer Inc. | Achievement of first commercial sale milestones | SB-525 and other products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  | $ 475,000,000.0 |  |  |  |  |  |  |  |  |
Pfizer SB-525 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 134,000,000.0 |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 55,000,000.0 |  |  |  |  |
Agreement termination, term |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 15 years |  |  |  |  |
Research service fees |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 79,000,000.0 |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  | $ 0 |  |  |  |  |
Pfizer SB-525 | SB-525 and other products |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 55,000,000.0 |
Number of milestones achieved | milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 2 |
Number of products approved | product |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |  |
Number of milestones included in transaction price | royaltyFee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |  |
Pfizer SB-525 | Recognition of license and stand-ready fee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Cumulative milestone achieved |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 300,000 |  | 2,500,000 |  |  |  |
Pfizer SB-525 | Milestone achievement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Cumulative milestone achieved |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 100,000 |  | 1,100,000 |  |  |  |
Sanofi |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  | $ 20,000,000.0 |  |  |  |  |  |  |  |
Agreement termination, term |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 180 days |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  |  |  |  | 6,000,000.0 |  | $ 6,000,000.0 |  |  |  | $ 6,000,000.0 |
Number of products approved | product |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 957,000 | 1,792,000 | $ 2,017,000 | 2,291,000 |  |  |  |
Number of research programs | program |  |  |  |  |  |  |  |  |  |  |  |  | 2 |  |  |  |  |  |  |  |
Sanofi | Recognition of license and stand-ready fee |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 206,000 | 380,000 | 433,000 | (349,000) |  |  |  |
Sanofi | Milestone achievement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Revenues |  |  |  |  |  |  |  |  |  |  |  |  |  | 138,000 | $ 257,000 | 292,000 | $ (235,000) |  |  |  |
Sanofi | Collaboration and license agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  | $ 276,300,000 |  |  |  |  |  |  |  |
Collaborative arrangement transaction price |  |  |  |  |  |  |  |  |  |  |  |  | 93,300,000 |  |  |  |  |  |  |  |
Collaborative arrangement estimated reimbursable service costs |  |  |  |  |  |  |  |  |  |  |  |  | 59,800,000 |  |  |  |  |  |  |  |
Deferred revenue |  |  |  |  |  |  |  |  |  |  |  |  |  | 500,000 |  | $ 500,000 |  | $ 1,200,000 |  | 500,000 |
Revenues under agreement |  |  |  |  |  |  |  |  |  |  |  |  | 20,000,000.0 |  |  |  |  |  |  |  |
Number of milestones included in transaction price | milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 0 |  |  |  |  |
Cumulative milestone achieved |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 7,400,000 |  |  |  |  |
Sanofi | Collaboration and license agreement | Milestone two |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  |  |  |  | 7,500,000 |  | 7,500,000 |  |  |  | 7,500,000 | $ 7,500,000
Milestone revenue reversal |  |  |  |  |  |  |  |  |  |  |  |  |  | 100,000 |  | 200,000 |  |  |  |  |
Sanofi | Collaboration and license agreement | Milestone three |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone revenue receivable |  |  |  |  |  |  |  |  |  |  |  |  |  | 13,500,000 |  | 13,500,000 |  |  |  | $ 13,500,000 |
Sanofi | Collaboration and license agreement | ST-40 Beta Thalassemia Phase 1 clinical trial milestone |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Milestone payments received |  |  |  |  |  |  |  | $ 6,000,000.0 |  |  |  |  |  |  |  |  |  |  |  |  |
Cumulative milestone achieved |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | 5,900,000 |  |  |  |  |
Milestone revenue reversal |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 100,000 |  | $ 100,000 |  |  |  |  |
Sanofi | Collaboration and license agreement | Achievement of specified clinical development And regulatory milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  | 115,800,000 |  |  |  |  |  |  |  |
Sanofi | Collaboration and license agreement | Achievement of specified sales milestones |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Research and Development Arrangement, Contract to Perform for Others [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Development and sales-based milestone payments to be received |  |  |  |  |  |  |  |  |  |  |  |  | $ 160,500,000 |  |  |  |  |  |  |  |

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