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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Deficit
Accumulated Other Comprehensive Income (Loss)
Non- Controlling Interest
Beginning Balances (in shares) at Dec. 31, 2019   115,972,708        
Beginning Balances at Dec. 31, 2019 $ 432,739 $ 1,160 $ 1,090,828 $ (656,985) $ (2,449) $ 185
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock (in shares)   24,420,157        
Issuance of common stock 142,526 $ 244 142,282      
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   409,107        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 852 $ 4 848      
Issuance of common stock under employee stock purchase plan (in shares)   171,305        
Issuance of common stock under employee stock purchase plan 1,187 $ 2 1,185      
Stock-based compensation 12,384   12,384      
Acquisition of additional shares of Sangamo France (339)         (339)
Foreign currency translation adjustment 150       150  
Net unrealized gain on marketable securities, net of tax 646       646  
Buy-out of non-controlling interest 0          
Net loss (78,939)     (78,842)   (97)
Ending Balances (in shares) at Jun. 30, 2020   140,973,277        
Ending Balances at Jun. 30, 2020 511,206 $ 1,410 1,247,527 (735,827) (1,653) (251)
Beginning Balances (in shares) at Mar. 31, 2020   116,278,553        
Beginning Balances at Mar. 31, 2020 394,415 $ 1,163 1,096,854 (699,898) (3,828) 124
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock (in shares)   24,420,157        
Issuance of common stock 142,526 $ 244 142,282      
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   103,262        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 443 $ 1 442      
Issuance of common stock under employee stock purchase plan (in shares)   171,305        
Issuance of common stock under employee stock purchase plan 1,187 $ 2 1,185      
Stock-based compensation 6,764   6,764      
Acquisition of additional shares of Sangamo France (339)         (339)
Foreign currency translation adjustment 1,783       1,783  
Net unrealized gain on marketable securities, net of tax 392       392  
Net loss (35,965)     (35,929)   (36)
Ending Balances (in shares) at Jun. 30, 2020   140,973,277        
Ending Balances at Jun. 30, 2020 511,206 $ 1,410 1,247,527 (735,827) (1,653) (251)
Beginning Balances (in shares) at Dec. 31, 2020   142,063,203        
Beginning Balances at Dec. 31, 2020 497,366 $ 1,421 1,269,375 (777,981) 5,419 (868)
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock (in shares)   1,805,227        
Issuance of common stock 24,732 $ 18 24,714      
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   993,901        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 977 $ 10 967      
Issuance of common stock under employee stock purchase plan (in shares)   244,570        
Issuance of common stock under employee stock purchase plan 2,054 $ 2 2,052      
Stock-based compensation 17,007   17,007      
Acquisition of additional shares of Sangamo France (134)   (70)     (64)
Foreign currency translation adjustment (3,441)       (3,441)  
Net unrealized gain on marketable securities, net of tax 9       9  
Buy-out of non-controlling interest 943   (943)     943
Net loss (93,113)     (93,102)   (11)
Ending Balances (in shares) at Jun. 30, 2021   145,106,901        
Ending Balances at Jun. 30, 2021 445,457 $ 1,451 1,313,102 (871,083) 1,987 0
Beginning Balances (in shares) at Mar. 31, 2021   143,713,765        
Beginning Balances at Mar. 31, 2021 469,417 $ 1,437 1,292,118 (823,914) 714 (938)
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock (in shares)   770,465        
Issuance of common stock 9,081 $ 8 9,073      
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   378,101        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax 1,393 $ 4 1,389      
Issuance of common stock under employee stock purchase plan (in shares)   244,570        
Issuance of common stock under employee stock purchase plan 2,054 $ 2 2,052      
Stock-based compensation 9,483   9,483      
Acquisition of additional shares of Sangamo France (70)   (70)      
Foreign currency translation adjustment 1,308       1,308  
Net unrealized gain on marketable securities, net of tax (35)       (35)  
Buy-out of non-controlling interest     (943)     943
Net loss (47,174)     (47,169)   (5)
Ending Balances (in shares) at Jun. 30, 2021   145,106,901        
Ending Balances at Jun. 30, 2021 $ 445,457 $ 1,451 $ 1,313,102 $ (871,083) $ 1,987 $ 0