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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($) $ in Thousands | 3 Months Ended
--- | ---
Mar. 31, 2022 | Mar. 31, 2021
Operating Activities: |  |
Net loss | $ (43,977) | $ (45,939)
Adjustments to reconcile net loss to net cash used in operating activities: |  |
Depreciation and amortization | 2,832 | 1,875
Amortization of premium on marketable securities | 164 | 824
Amortization and other changes in operating lease right-of-use assets | 2,106 | 2,023
Loss on free shares | 0 | 27
Stock-based compensation | 7,691 | 7,524
Net changes in operating assets and liabilities: |  |
Accounts receivable | (925) | (1,522)
Prepaid expenses and other assets | (248) | (1,680)
Accounts payable and other accrued liabilities | 3,609 | (3,829)
Accrued compensation and employee benefits | (8,270) | (7,561)
Deferred revenues | (21,004) | (20,047)
Long-term portion of lease liabilities | (972) | (1,045)
Other non-current liabilities | 19 | 800
Net cash used in operating activities | (58,975) | (68,550)
Investing Activities: |  |
Purchases of marketable securities | (65,756) | (97,935)
Maturities of marketable securities | 70,300 | 144,369
Sales of marketable securities | 0 | 6,870
Purchases of property and equipment | (2,839) | (7,950)
Purchase of additional Sangamo France shares | 0 | (65)
Net cash provided by investing activities | 1,705 | 45,289
Financing Activities: |  |
Taxes paid related to net share settlement of equity awards | (1,641) | (2,234)
Proceeds from exercise of stock options and restricted stock units | 74 | 1,818
Net cash (used in) provided by financing activities | (1,567) | 15,235
Effect of exchange rate changes on cash, cash equivalents, and restricted cash | 222 | (328)
Net decrease in cash, cash equivalents, and restricted cash | (58,615) | (8,354)
Cash, cash equivalents, and restricted cash, beginning of period | 180,372 | 132,829
Cash, cash equivalents, and restricted cash, end of period | 121,757 | 124,475
Supplemental cash flow disclosures: |  |
Property and equipment included in unpaid liabilities | 2,257 | 2,953
Right-of-use assets obtained in exchange for lease obligations | 0 | 1,356
At the Market Offering |  |
Financing Activities: |  |
Proceeds from at-the-market offering, net of offering expenses | $ 0 | $ 15,651

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