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CONDENSED CONSOLIDATED STATEMENTS OF STOCKHOLDERS' EQUITY - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Deficit
Accumulated Other Comprehensive (Loss) Income
Non- Controlling Interest
Beginning balance (in shares) at Dec. 31, 2020   142,063,203        
Beginning balance at Dec. 31, 2020 $ 497,366 $ 1,421 $ 1,269,375 $ (777,981) $ 5,419 $ (868)
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock in connection with at-the-market offering, net of offering expenses (in shares)   1,034,762        
Issuance of common stock in connection with at-the-market offering, net of offering expenses 15,651 $ 10 15,641      
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   615,800        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (416) $ 6 (422)      
Stock-based compensation 7,524   7,524      
Acquisition of additional shares of Sangamo France (64)         (64)
Foreign currency translation adjustment (4,749)       (4,749)  
Net pension gains 0          
Net unrealized loss on marketable securities, net of tax 44       44  
Net loss (45,939)     (45,933)   (6)
Ending balance (in shares) at Mar. 31, 2021   143,713,765        
Ending balance at Mar. 31, 2021 469,417 $ 1,437 1,292,118 (823,914) 714 $ (938)
Beginning balance (in shares) at Dec. 31, 2021   145,921,530        
Beginning balance at Dec. 31, 2021 375,343 $ 1,459 1,334,138 (956,267) (3,987)  
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (in shares)   743,230        
Issuance of common stock upon exercise of stock options and in connection with restricted stock units, net of tax (1,567) $ 8 (1,575)      
Stock-based compensation 7,691   7,691      
Foreign currency translation adjustment (1,904)       (1,904)  
Net pension gains 19       19  
Net unrealized loss on marketable securities, net of tax (1,083)       (1,083)  
Net loss (43,977)     (43,977)    
Ending balance (in shares) at Mar. 31, 2022   146,664,760        
Ending balance at Mar. 31, 2022 $ 334,522 $ 1,467 $ 1,340,254 $ (1,000,244) $ (6,955)