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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2022
Mar. 31, 2021
Operating Activities:    
Net loss $ (43,977) $ (45,939)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 2,832 1,875
Amortization of premium on marketable securities 164 824
Amortization and other changes in operating lease right-of-use assets 2,106 2,023
Loss on free shares 0 27
Stock-based compensation 7,691 7,524
Net changes in operating assets and liabilities:    
Accounts receivable (925) (1,522)
Prepaid expenses and other assets (248) (1,680)
Accounts payable and other accrued liabilities 3,609 (3,829)
Accrued compensation and employee benefits (8,270) (7,561)
Deferred revenues (21,004) (20,047)
Long-term portion of lease liabilities (972) (1,045)
Other non-current liabilities 19 800
Net cash used in operating activities (58,975) (68,550)
Investing Activities:    
Purchases of marketable securities (65,756) (97,935)
Maturities of marketable securities 70,300 144,369
Sales of marketable securities 0 6,870
Purchases of property and equipment (2,839) (7,950)
Purchase of additional Sangamo France shares 0 (65)
Net cash provided by investing activities 1,705 45,289
Financing Activities:    
Taxes paid related to net share settlement of equity awards (1,641) (2,234)
Proceeds from exercise of stock options and restricted stock units 74 1,818
Net cash (used in) provided by financing activities (1,567) 15,235
Effect of exchange rate changes on cash, cash equivalents, and restricted cash 222 (328)
Net decrease in cash, cash equivalents, and restricted cash (58,615) (8,354)
Cash, cash equivalents, and restricted cash, beginning of period 180,372 132,829
Cash, cash equivalents, and restricted cash, end of period 121,757 124,475
Supplemental cash flow disclosures:    
Property and equipment included in unpaid liabilities 2,257 2,953
Right-of-use assets obtained in exchange for lease obligations 0 1,356
At the Market Offering    
Financing Activities:    
Proceeds from at-the-market offering, net of offering expenses $ 0 $ 15,651