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CONDENSED CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 9 Months Ended
Sep. 30, 2022
Sep. 30, 2021
Operating Activities:    
Net loss $ (140,305) $ (140,801)
Adjustments to reconcile net loss to net cash used in operating activities:    
Depreciation and amortization 8,765 6,675
Amortization of (discount) premium on marketable securities (324) 2,394
Amortization and other changes in operating lease right-of-use assets 6,357 6,104
Gain on free shares 0 (18)
Stock-based compensation 23,402 24,880
Gain on disposal of property and equipment 0 (30)
Net changes in operating assets and liabilities:    
Interest receivable (175) 415
Accounts receivable 1,491 (2,743)
Prepaid expenses and other assets (5,848) (6,467)
Accounts payable and other accrued liabilities 9,443 (4,699)
Accrued compensation and employee benefits (1,225) (1,932)
Deferred revenues (64,956) (62,292)
Lease liabilities (3,298) (3,227)
Other non-current liabilities 95 1,204
Net cash used in operating activities (166,578) (180,537)
Investing Activities:    
Purchases of marketable securities (225,621) (300,387)
Maturities of marketable securities 254,990 509,620
Sales of marketable securities 0 6,870
Purchases of property and equipment (12,697) (20,420)
Purchase of additional Sangamo France shares 0 (119)
Net cash provided by investing activities 16,672 195,564
Financing Activities:    
Taxes paid related to net share settlement of equity awards (1,962) (2,988)
Proceeds from exercise of stock options 124 4,582
Net cash provided by financing activities 65,125 30,746
Effect of exchange rate changes on cash, cash equivalents, and restricted cash 479 (148)
Net (decrease) increase in cash, cash equivalents, and restricted cash (84,302) 45,625
Cash, cash equivalents, and restricted cash, beginning of period 180,372 132,829
Cash, cash equivalents, and restricted cash, end of period 96,070 178,454
Supplemental cash flow disclosures:    
Property and equipment included in unpaid liabilities 3,927 650
Right-of-use assets obtained in exchange for lease obligations 0 1,349
Tenant improvement allowance included in contra-lease liability 1,531 0
Buy-out of non-controlling interest 0 943
At the Market Offering    
Financing Activities:    
Proceeds from at-the market offering/ issuance of common stock 65,848 27,099
Employee Stock    
Financing Activities:    
Proceeds from at-the market offering/ issuance of common stock $ 1,115 $ 2,053