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CONSOLIDATED STATEMENTS OF STOCKHOLDERS’ EQUITY - USD ($)
$ in Thousands
Total
Common Stock
Additional Paid-in Capital
Accumulated Deficit
Accumulated Other Comprehensive (Loss) Income
Non- Controlling Interest
Beginning balance (in shares) at Dec. 31, 2020   142,063,203        
Beginning balance at Dec. 31, 2020 $ 497,366 $ 1,421 $ 1,269,375 $ (777,981) $ 5,419 $ (868)
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock in at-the-market offering, net of offering expenses (in shares)   2,007,932        
Issuance of common stock in at-the-market offering, net of offering expenses 27,099 $ 20 27,079      
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares)   1,417,288        
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax $ 2,389 $ 14 2,375      
Issuance of common stock under employee stock purchase plan (in shares) 433,107 433,107        
Issuance of common stock under employee stock purchase plan $ 3,370 $ 4 3,366      
Stock-based compensation 32,956   32,956      
Acquisition of additional shares of Sangamo France (134)   (70)     (64)
Foreign currency translation adjustment (8,351)       (8,351)  
Net pension gain (loss) (716)       (716)  
Net unrealized gain (loss) on marketable securities, net of tax (339)       (339)  
Buy-out of non-controlling interest 943   (943)     943
Net loss (178,297)     (178,286)   (11)
Ending balance (in shares) at Dec. 31, 2021   145,921,530        
Ending balance at Dec. 31, 2021 375,343 $ 1,459 1,334,138 (956,267) (3,987) $ 0
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock in at-the-market offering, net of offering expenses (in shares)   19,300,743        
Issuance of common stock in at-the-market offering, net of offering expenses 84,869 $ 193 84,676      
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares)   994,097        
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax $ (1,980) $ 10 (1,990)      
Issuance of common stock under employee stock purchase plan (in shares) 576,950 576,950        
Issuance of common stock under employee stock purchase plan $ 1,771 $ 6 1,765      
Stock-based compensation 31,650   31,650      
Foreign currency translation adjustment (4,606)       (4,606)  
Net pension gain (loss) 786       786  
Net unrealized gain (loss) on marketable securities, net of tax (597)       (597)  
Buy-out of non-controlling interest 0          
Net loss $ (192,278)     (192,278)    
Ending balance (in shares) at Dec. 31, 2022 166,793,320 166,793,320        
Ending balance at Dec. 31, 2022 $ 294,958 $ 1,668 1,450,239 (1,148,545) (8,404)  
Increase (Decrease) in Stockholders' Equity [Roll Forward]            
Issuance of common stock in at-the-market offering, net of offering expenses (in shares)   8,249,261        
Issuance of common stock in at-the-market offering, net of offering expenses 15,106 $ 82 15,024      
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares)   1,744,118        
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax $ (1,453) $ 18 (1,471)      
Issuance of common stock under employee stock purchase plan (in shares) 1,346,849 1,346,849        
Issuance of common stock under employee stock purchase plan $ 935 $ 13 922      
Stock-based compensation 27,363   27,363      
Foreign currency translation adjustment 2,848       2,848  
Net pension gain (loss) (163)       (163)  
Net unrealized gain (loss) on marketable securities, net of tax 1,124       1,124  
Buy-out of non-controlling interest 0          
Net loss $ (257,831)     (257,831)    
Ending balance (in shares) at Dec. 31, 2023 178,133,548 178,133,548        
Ending balance at Dec. 31, 2023 $ 82,887 $ 1,781 $ 1,492,077 $ (1,406,376) $ (4,595)