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CONSOLIDATED STATEMENTS OF STOCKHOLDERS’ EQUITY - USD ($)
$ in Thousands
Total
At-The-Market Offering
Stock Issuances Excluding At-The-Market Offering
Common Stock
Common Stock
At-The-Market Offering
Common Stock
Stock Issuances Excluding At-The-Market Offering
Additional Paid-in Capital
Additional Paid-in Capital
At-The-Market Offering
Additional Paid-in Capital
Stock Issuances Excluding At-The-Market Offering
Accumulated Deficit
Accumulated Other Comprehensive Loss
Beginning balance (in shares) at Dec. 31, 2022       166,793,320              
Beginning balance at Dec. 31, 2022 $ 294,958     $ 1,668     $ 1,450,239     $ (1,148,545) $ (8,404)
Increase (Decrease) in Stockholders' Equity [Roll Forward]                      
Issuance of common stock in at-the-market offering, net of offering expenses (in shares)       8,249,261              
Issuance of common stock in at-the-market offering, net of offering expenses 15,106     $ 82     15,024        
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares)       1,744,118              
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax $ (1,453)     $ 18     (1,471)        
Issuance of common stock under employee stock purchase plan (in shares) 1,346,849     1,346,849              
Issuance of common stock under employee stock purchase plan $ 935     $ 13     922        
Stock-based compensation 27,363           27,363        
Foreign currency translation adjustment 2,848                   2,848
Net pension gain (loss) (163)                   (163)
Net unrealized loss on marketable securities, net of tax 1,124                   1,124
Net loss $ (257,831)                 (257,831)  
Ending balance (in shares) at Dec. 31, 2023 178,133,548     178,133,548              
Ending balance at Dec. 31, 2023 $ 82,887     $ 1,781     1,492,077     (1,406,376) (4,595)
Increase (Decrease) in Stockholders' Equity [Roll Forward]                      
Issuance of common stock in at-the-market offering, net of offering expenses (in shares)         3,638,740 24,761,905          
Issuance of common stock in at-the-market offering, net of offering expenses   $ 7,132 $ 21,788   $ 36 $ 248   $ 7,096 $ 21,540    
Issuance of common stock upon exercise of pre-funded warrants (in shares)       3,809,523              
Issuance of common stock upon exercise of pre-funded warrants 71     $ 38     33        
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax (in shares)       1,746,271              
Issuance of common stock upon exercise of stock options and vesting of restricted stock units, net of tax $ (904)     $ 17     (921)        
Issuance of common stock under employee stock purchase plan (in shares) 747,692     747,692              
Issuance of common stock under employee stock purchase plan $ 290     $ 8     282        
Stock-based compensation 12,382           12,382        
Foreign currency translation adjustment (2,943)                   (2,943)
Net pension gain (loss) 241                   241
Net unrealized loss on marketable securities, net of tax (233)                   (233)
Net loss $ (97,941)                 (97,941)  
Ending balance (in shares) at Dec. 31, 2024 212,837,679     212,837,679              
Ending balance at Dec. 31, 2024 $ 22,770     $ 2,128     $ 1,532,489     $ (1,504,317) $ (7,530)