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CONSOLIDATED STATEMENTS OF CASH FLOWS - USD ($)
$ in Thousands
 12 Months Ended
Dec. 31, 2024
Dec. 31, 2023
Operating Activities:    
Net loss $ (97,941) $ (257,831)
Adjustments to reconcile net loss to net cash used in operating activities:    
Impairment of long-lived assets 5,521 65,528
Depreciation and amortization 5,107 15,065
Accretion of discount and impairment on marketable securities (273) (2,329)
Amortization in operating lease right-of-use assets 4,254 7,127
Stock-based compensation 12,382 27,363
Gain on sale of assets classified as held for sale (1,026) 0
Other 330 1,119
Impairment of goodwill and indefinite-lived intangible assets 0 89,485
Deferred income tax benefit 0 (6,335)
Net changes in operating assets and liabilities:    
Accounts receivable 397 2,755
Refundable research income tax credits 556 (3,731)
Prepaid expenses and other assets 4,384 10,256
Accounts payable and other accrued liabilities (14,169) 5,589
Accrued compensation and employee benefits 5,743 (12,690)
Deferred revenues 13,430 (161,156)
Lease liabilities (5,579) (5,037)
Other non-current liabilities (255) (20)
Net cash used in operating activities (67,139) (224,842)
Investing Activities:    
Purchases of marketable securities 0 (59,551)
Maturities of marketable securities 1,110 214,500
Proceeds from sale of marketable securities 34,730 19,737
Proceeds from sale of assets classified as held for sale 1,951 0
Purchases of property and equipment (267) (21,155)
Net cash provided by investing activities 37,524 153,531
Financing Activities:    
Proceeds from at-the-market offering, net of offering expenses 7,132 15,106
Proceeds from issuance of common stock, net of offering expenses 21,859 0
Taxes paid related to net share settlement of equity awards (904) (1,453)
Proceeds from issuance of common stock under employee stock purchase plan 290 935
Net cash provided by financing activities 28,377 14,588
Effect of exchange rate changes on cash and cash equivalents and restricted cash (2,048) 1,483
Net decrease in cash, cash equivalents and restricted cash (3,286) (55,240)
Cash, cash equivalents and restricted cash, beginning of period 46,704 101,944
Cash, cash equivalents and restricted cash, end of period 43,418 46,704
Supplemental cash flow disclosures:    
Property and equipment included in unpaid liabilities $ 236 $ 447